Difference between revisions of "Grindelia hirsutula"

Hooker & Arnott

Bot. Beechey Voy., 147. 1833.

Synonyms: Grindelia acutifolia Steyermark Grindelia camporum Greene Grindelia camporum var. bracteosa (J. T. Howell) M. A. Lane Grindelia camporum var. davyi (Jepson) Steyermark Grindelia camporum var. parviflora Steyermark Grindelia columbiana Hooker & Arnott Grindelia fastigiata Greene Grindelia hallii de Candolle Grindelia hirsutula var. davyi (Jepson) M. A. Lane Grindelia hirsutula var. hallii (Steyermark) M. A. Lane Grindelia hirsutula var. maritima (Greene) M. A. Lane Grindelia hirsutula subsp. rubricaulis (de Candolle) D. D. Keck Grindelia humilis Greene Grindelia inornata A. Nelson Grindelia inornata var. angusta Steyermark Grindelia integrifolia var. macrophylla (Greene) Cronquist Grindelia latifolia Nuttall Grindelia latifolia subsp. platyphylla (Greene) D. D. Keck Grindelia macrophylla (Pursh) Dunal Grindelia maritima de Candolle Grindelia nana unknown Grindelia nana var. altissima Steyermark Grindelia nana subsp. columbiana Piper Grindelia nana var. integerrima (Rydberg) Steyermark Grindelia nana var. integrifolia Nuttall Grindelia paludosa unknown Grindelia perennis unknown Grindelia procera unknown Grindelia revoluta unknown Grindelia robusta unknown Grindelia robusta var. bracteosa (J. T. Howell) D. D. Keck Grindelia robusta var. davyi Jepson Grindelia robusta var. platyphylla Greene Grindelia robusta var. rigida A. Gray Grindelia rubricaulis unknown Grindelia rubricaulis var. maritima Greene Grindelia squarrosa var. integrifolia (Nuttall) B. Boivin Grindelia squarrosa var. quasiperennis Lunell Grindelia stricta unknown Grindelia stricta var. angustifolia (A. Gray) M. A. Lane Grindelia stricta subsp. blakei (Steyermark) D. D. Keck Grindelia stricta var. macrophylla (Greene) Steyermark Grindelia stricta var. platyphylla (Greene) M. A. Lane Grindelia stricta subsp. venulosa (Jepson) D. D. Keck
Treatment appears in FNA Volume 20. Treatment on page 432. Mentioned on page 425, 427, 428, 429, 431, 433, 434, 435.
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|name=Grindelia acutifolia
 
|name=Grindelia acutifolia
 
|authority=Steyermark
 
|authority=Steyermark
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia camporum
 
|name=Grindelia camporum
 
|authority=Greene
 
|authority=Greene
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia camporum var. bracteosa
 
|name=Grindelia camporum var. bracteosa
 
|authority=(J. T. Howell) M. A. Lane
 
|authority=(J. T. Howell) M. A. Lane
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia camporum var. davyi
 
|name=Grindelia camporum var. davyi
 
|authority=(Jepson) Steyermark
 
|authority=(Jepson) Steyermark
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia camporum var. parviflora
 
|name=Grindelia camporum var. parviflora
 
|authority=Steyermark
 
|authority=Steyermark
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia columbiana
 
|name=Grindelia columbiana
 
|authority=Hooker & Arnott
 
|authority=Hooker & Arnott
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia fastigiata
 
|name=Grindelia fastigiata
 
|authority=Greene
 
|authority=Greene
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia hallii
 
|name=Grindelia hallii
 
|authority=de Candolle
 
|authority=de Candolle
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia hirsutula var. davyi
 
|name=Grindelia hirsutula var. davyi
 
|authority=(Jepson) M. A. Lane
 
|authority=(Jepson) M. A. Lane
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia hirsutula var. hallii
 
|name=Grindelia hirsutula var. hallii
 
|authority=(Steyermark) M. A. Lane
 
|authority=(Steyermark) M. A. Lane
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia hirsutula var. maritima
 
|name=Grindelia hirsutula var. maritima
 
|authority=(Greene) M. A. Lane
 
|authority=(Greene) M. A. Lane
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia hirsutula subsp. rubricaulis
 
|name=Grindelia hirsutula subsp. rubricaulis
 
|authority=(de Candolle) D. D. Keck
 
|authority=(de Candolle) D. D. Keck
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia humilis
 
|name=Grindelia humilis
 
|authority=Greene
 
|authority=Greene
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia inornata
 
|name=Grindelia inornata
 
|authority=A. Nelson
 
|authority=A. Nelson
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia inornata var. angusta
 
|name=Grindelia inornata var. angusta
 
|authority=Steyermark
 
|authority=Steyermark
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia integrifolia var. macrophylla
 
|name=Grindelia integrifolia var. macrophylla
 
|authority=(Greene) Cronquist
 
|authority=(Greene) Cronquist
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia latifolia
 
|name=Grindelia latifolia
 
|authority=Nuttall
 
|authority=Nuttall
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia latifolia subsp. platyphylla
 
|name=Grindelia latifolia subsp. platyphylla
 
|authority=(Greene) D. D. Keck
 
|authority=(Greene) D. D. Keck
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia macrophylla
 
|name=Grindelia macrophylla
 
|authority=(Pursh) Dunal
 
|authority=(Pursh) Dunal
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia maritima
 
|name=Grindelia maritima
 
|authority=de Candolle
 
|authority=de Candolle
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia nana
 
|name=Grindelia nana
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia nana var. altissima
 
|name=Grindelia nana var. altissima
 
|authority=Steyermark
 
|authority=Steyermark
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia nana subsp. columbiana
 
|name=Grindelia nana subsp. columbiana
 
|authority=Piper
 
|authority=Piper
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia nana var. integerrima
 
|name=Grindelia nana var. integerrima
 
|authority=(Rydberg) Steyermark
 
|authority=(Rydberg) Steyermark
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia nana var. integrifolia
 
|name=Grindelia nana var. integrifolia
 
|authority=Nuttall
 
|authority=Nuttall
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia paludosa
 
|name=Grindelia paludosa
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia perennis
 
|name=Grindelia perennis
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia procera
 
|name=Grindelia procera
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia revoluta
 
|name=Grindelia revoluta
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia robusta
 
|name=Grindelia robusta
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia robusta var. bracteosa
 
|name=Grindelia robusta var. bracteosa
 
|authority=(J. T. Howell) D. D. Keck
 
|authority=(J. T. Howell) D. D. Keck
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia robusta var. davyi
 
|name=Grindelia robusta var. davyi
 
|authority=Jepson
 
|authority=Jepson
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia robusta var. platyphylla
 
|name=Grindelia robusta var. platyphylla
 
|authority=Greene
 
|authority=Greene
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia robusta var. rigida
 
|name=Grindelia robusta var. rigida
 
|authority=A. Gray
 
|authority=A. Gray
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia rubricaulis
 
|name=Grindelia rubricaulis
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia rubricaulis var. maritima
 
|name=Grindelia rubricaulis var. maritima
 
|authority=Greene
 
|authority=Greene
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia squarrosa var. integrifolia
 
|name=Grindelia squarrosa var. integrifolia
 
|authority=(Nuttall) B. Boivin
 
|authority=(Nuttall) B. Boivin
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia squarrosa var. quasiperennis
 
|name=Grindelia squarrosa var. quasiperennis
 
|authority=Lunell
 
|authority=Lunell
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia stricta
 
|name=Grindelia stricta
 
|authority=unknown
 
|authority=unknown
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia stricta var. angustifolia
 
|name=Grindelia stricta var. angustifolia
 
|authority=(A. Gray) M. A. Lane
 
|authority=(A. Gray) M. A. Lane
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia stricta subsp. blakei
 
|name=Grindelia stricta subsp. blakei
 
|authority=(Steyermark) D. D. Keck
 
|authority=(Steyermark) D. D. Keck
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia stricta var. macrophylla
 
|name=Grindelia stricta var. macrophylla
 
|authority=(Greene) Steyermark
 
|authority=(Greene) Steyermark
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia stricta var. platyphylla
 
|name=Grindelia stricta var. platyphylla
 
|authority=(Greene) M. A. Lane
 
|authority=(Greene) M. A. Lane
}}{{Treatment/ID/Synonym
+
}} {{Treatment/ID/Synonym
 
|name=Grindelia stricta subsp. venulosa
 
|name=Grindelia stricta subsp. venulosa
 
|authority=(Jepson) D. D. Keck
 
|authority=(Jepson) D. D. Keck
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|elevation=0–2800 m
 
|elevation=0–2800 m
 
|distribution=Alta.;B.C.;Man.;N.W.T.;Ont.;Que.;Sask.;Alaska;Ariz.;Calif.;Colo.;Idaho;Ill.;Mich.;Minn.;Mo.;Mont.;Nebr.;Nev.;N.Mex.;N.Y.;N.Dak.;Oreg.;Pa.;S.Dak.;Utah;Wash.;Wyo.;introduced in Mexico (Yucatan).
 
|distribution=Alta.;B.C.;Man.;N.W.T.;Ont.;Que.;Sask.;Alaska;Ariz.;Calif.;Colo.;Idaho;Ill.;Mich.;Minn.;Mo.;Mont.;Nebr.;Nev.;N.Mex.;N.Y.;N.Dak.;Oreg.;Pa.;S.Dak.;Utah;Wash.;Wyo.;introduced in Mexico (Yucatan).
|discussion=<p>Occurrence of Grindelia hirsutula in Alaska was not verified for this treatment. As circumscribed here, Grindelia hirsutula includes 30 or more reputedly distinct, local, regional, or ecotypic facies that have been named at species or infraspecific rank. Locally, such facies are easily recognized; in a broad view, they intergrade with other facies and are parts of a heterogeneous continuum. Taxonomies that have attempted to recognize the facies as distinct “taxa” have led to almost as many specimens determined as “intermediate” as are assigned to the named “taxa.”</p><!--
+
|discussion=<p>Occurrence of <i>Grindelia hirsutula</i> in Alaska was not verified for this treatment. As circumscribed here, <i>Grindelia hirsutula</i> includes 30 or more reputedly distinct, local, regional, or ecotypic facies that have been named at species or infraspecific rank. Locally, such facies are easily recognized; in a broad view, they intergrade with other facies and are parts of a heterogeneous continuum. Taxonomies that have attempted to recognize the facies as distinct “taxa” have led to almost as many specimens determined as “intermediate” as are assigned to the named “taxa.”</p><!--
--><p>Following, alphabetically by epithet, we have summarized names, diagnostic traits (mostly as given in regional and/or state floras), and distributions for “taxa” treated as distinct in some taxonomies and included within Grindelia hirsutula here:</p><!--
+
--><p>Following, alphabetically by epithet, we have summarized names, diagnostic traits (mostly as given in regional and/or state floras), and distributions for “taxa” treated as distinct in some taxonomies and included within <i>Grindelia hirsutula</i> here:</p><!--
--><p>acutifolia: Grindelia acutifolia; cauline leaf blades firmly membranous, lengths 2.5–5 times widths, phyllary apices looped to hooked, pappi of 2–3, subulate scales; southeastern Colorado, northeastern New Mexico Hybrids between G. acutifolia and G. squarrosa (as G. nuda) have been recorded from the Colorado side of Raton Pass.</p><!--
+
--><p>acutifolia: <i>Grindelia</i> acutifolia; cauline leaf blades firmly membranous, lengths 2.5–5 times widths, phyllary apices looped to hooked, pappi of 2–3, subulate scales; southeastern Colorado, northeastern New Mexico Hybrids between G. acutifolia and <i>G. squarrosa</i> (as G. nuda) have been recorded from the Colorado side of Raton Pass.</p><!--
--><p>altissima: Grindelia nana var. altissima; distal cauline leaves smaller and more scattered than proximal, blades about as wide at bases as at mid points, heads relatively few, heights of involucres less than diams.; mostly inner coastal ranges, northern California, southern Oregon. M. A. Lane (1993b) considered altissima to apply to hybrids between davyi and nana.</p><!--
+
--><p>altissima: <i>Grindelia</i> nana <i></i>var.<i> altissima</i>; distal cauline leaves smaller and more scattered than proximal, blades about as wide at bases as at mid points, heads relatively few, heights of involucres less than diams.; mostly inner coastal ranges, northern California, southern Oregon. M. A. Lane (1993b) considered altissima to apply to hybrids between davyi and nana.</p><!--
--><p>angusta: Grindelia inornata var. angusta; cauline leaf blades oblong to oblong-lanceolate, lengths 3+ times widths, faces inconspicuously gland-dotted, ray florets 0, cypselae stramineous to pale brown; central Colorado.</p><!--
+
--><p>angusta: <i>Grindelia</i> inornata var. angusta; cauline leaf blades oblong to oblong-lanceolate, lengths 3+ times widths, faces inconspicuously gland-dotted, ray florets 0, cypselae stramineous to pale brown; central Colorado.</p><!--
--><p>angustifolia: Grindelia stricta var. angustifolia; see humilis.</p><!--
+
--><p>angustifolia: <i>Grindelia</i> stricta <i></i>var.<i> angustifolia</i>; see humilis.</p><!--
--><p>blakei: Grindelia stricta subsp. blakei; see humilis.</p><!--
+
--><p>blakei: <i>Grindelia</i> stricta subsp. blakei; see humilis.</p><!--
--><p>bracteosa: Grindelia bracteosa; G. camporum var. bracteosa; G. robusta var. bracteosa; stems usually whitish and resinous, lengths of cauline leaf blades 3–5 times widths, phyllary apices usually looped, ray florets 0 or 2–27; southern California. M. A. Lane (1993b) suggested that bracteosa may have derived from hybridization between hirsutula and G. squarrosa (var. serrulata). See also, robusta.</p><!--
+
--><p>bracteosa: <i>Grindelia</i> bracteosa; G. camporum <i></i>var.<i> bracteosa</i>; G. robusta <i></i>var.<i> bracteosa</i>; stems usually whitish and resinous, lengths of cauline leaf blades 3–5 times widths, phyllary apices usually looped, ray florets 0 or 2–27; southern California. M. A. Lane (1993b) suggested that bracteosa may have derived from hybridization between hirsutula and <i>G. squarrosa</i> (var. serrulata). See also, robusta.</p><!--
--><p>camporum: Grindelia camporum; stems usually whitish and resinous, cauline leaf blades ovate to lanceolate, involucres seldom subtended by bracts, phyllary apices recurved to straight, ray florets 32–39; mostly interior north, central California.</p><!--
+
--><p>camporum: <i>Grindelia</i> camporum; stems usually whitish and resinous, cauline leaf blades ovate to lanceolate, involucres seldom subtended by bracts, phyllary apices recurved to straight, ray florets 32–39; mostly interior north, central California.</p><!--
--><p>columbiana: Grindelia nana [unranked] columbiana; G. columbiana; ray florets 0, otherwise much like nana; Idaho, Oregon, Washington (mostly Columbia River drainage).</p><!--
+
--><p>columbiana: <i>Grindelia</i> nana [unranked] columbiana; G. columbiana; ray florets 0, otherwise much like nana; Idaho, Oregon, Washington (mostly Columbia River drainage).</p><!--
--><p>davyi: Grindelia camporum var. davyi; G. hirsutula var. davyi; G. robusta var. davyi; phyllary apices recurved to straight, acuminate, pappi usually ± equaling disc corollas; interior northern California. M. A. Lane (1993b) suggested that davyi may have derived from hybridization between camporum and hirsutula.</p><!--
+
--><p>davyi: <i>Grindelia</i> camporum <i></i>var.<i> davyi</i>; <i>G. hirsutula</i> <i></i>var.<i> davyi</i>; G. robusta <i></i>var.<i> davyi</i>; phyllary apices recurved to straight, acuminate, pappi usually ± equaling disc corollas; interior northern California. M. A. Lane (1993b) suggested that davyi may have derived from hybridization between camporum and hirsutula.</p><!--
--><p>fastigiata: Grindelia fastigiata; cauline leaf blades conspicuously gland-dotted, phyllary apices thick, coriaceous, ray florets 0, cypselae brown; western Colorado and eastern Utah. Hybrids between fastigiata and G. squarrosa have been recorded from Utah.</p><!--
+
--><p>fastigiata: <i>Grindelia</i> fastigiata; cauline leaf blades conspicuously gland-dotted, phyllary apices thick, coriaceous, ray florets 0, cypselae brown; western Colorado and eastern Utah. Hybrids between fastigiata and <i>G. squarrosa</i> have been recorded from Utah.</p><!--
--><p>hallii: Grindelia hallii; G. hirsutula var. hallii; involucres 8–12 mm diam., phyllary apices slightly recurved and attenuate to nearly straight and deltate, pappi shorter than disc corollas; interior southern California.</p><!--
+
--><p>hallii: <i>Grindelia</i> hallii; <i>G. hirsutula</i> <i></i>var.<i> hallii</i>; involucres 8–12 mm diam., phyllary apices slightly recurved and attenuate to nearly straight and deltate, pappi shorter than disc corollas; interior southern California.</p><!--
--><p>hirsutula: Grindelia hirsutula; stems usually green to reddish, ± hirsutulous to arachnose (at least distally), cauline leaf blades usually widest proximal to their midpoints, involucres 12–23 mm diam., often subtended by leaflike bracts, phyllary apices mostly recurved to straight, slightly to moderately resinous, ray florets usually 20–60, laminae 14–20 mm, cypselae usually reddish; mostly coastal ranges of California.</p><!--
+
--><p>hirsutula: <i>Grindelia hirsutula</i>; stems usually green to reddish, ± hirsutulous to arachnose (at least distally), cauline leaf blades usually widest proximal to their midpoints, involucres 12–23 mm diam., often subtended by leaflike bracts, phyllary apices mostly recurved to straight, slightly to moderately resinous, ray florets usually 20–60, laminae 14–20 mm, cypselae usually reddish; mostly coastal ranges of California.</p><!--
--><p>humilis: Grindelia humilis Hooker & Arnott; about the nomenclatural type of G. humilis, M. A. Lane (1992b) stated “...clearly conspecific with G. hirsutula” and she used the name G. stricta var. angustifolia for plants others had treated as G. humilis; as used by others, humilis has been applied to subshrubs 100–150 cm with red-brown stems and phyllary apices recurved to nearly straight from tidal lands around San Francisco and Tomales bays in northern California; similar plants but for apices of phyllaries often looped to hooked from around Humboldt Bay in northern California have been called blakei.</p><!--
+
--><p>humilis: <i>Grindelia</i> humilis Hooker & Arnott; about the nomenclatural type of G. humilis, M. A. Lane (1992b) stated “...clearly conspecific with <i>G. hirsutula</i>” and she used the name G. stricta <i></i>var.<i> angustifolia</i> for plants others had treated as G. humilis; as used by others, humilis has been applied to subshrubs 100–150 cm with red-brown stems and phyllary apices recurved to nearly straight from tidal lands around San Francisco and Tomales bays in northern California; similar plants but for apices of phyllaries often looped to hooked from around Humboldt Bay in northern California have been called blakei.</p><!--
--><p>inornata: Grindelia inornata; cauline leaf blades oblanceolate or ovate to oblong, lengths 1.5–3 times widths, faces inconspicuously gland-dotted, ray florets 0, cypselae stramineous to pale brown; central Colorado. Hybrids between inornata and G. subalpina have been recorded from Colorado.</p><!--
+
--><p>inornata: <i>Grindelia</i> inornata; cauline leaf blades oblanceolate or ovate to oblong, lengths 1.5–3 times widths, faces inconspicuously gland-dotted, ray florets 0, cypselae stramineous to pale brown; central Colorado. Hybrids between inornata and <i>G. subalpina</i> have been recorded from Colorado.</p><!--
--><p>integerrima: Grindelia nana var. integerrima; see integrifolia.</p><!--
+
--><p>integerrima: <i>Grindelia</i> nana var. integerrima; see integrifolia.</p><!--
--><p>integrifolia: Grindelia nana var. integrifolia Nuttall (not G. integrifolia de Candolle); G. squarrosa var. integrifolia; cauline leaf blades oblanceolate, margins subentire, phyllary apices (at least the outer) looped to hooked, pappi of curled, smooth to barbellulate, subulate scales scarcely longer than cypselae; Idaho, Montana, Nevada, Oregon, Utah, Washington.</p><!--
+
--><p>integrifolia: <i>Grindelia</i> nana <i></i>var.<i> integrifolia</i> Nuttall (not <i>G. integrifolia</i> de Candolle); <i>G. squarrosa</i> <i></i>var.<i> integrifolia</i>; cauline leaf blades oblanceolate, margins subentire, phyllary apices (at least the outer) looped to hooked, pappi of curled, smooth to barbellulate, subulate scales scarcely longer than cypselae; Idaho, Montana, <i>Nevada</i>, Oregon, Utah, Washington.</p><!--
--><p>latifolia: Grindelia latifolia; stems herbaceous, decumbent to prostrate, leaf blade apices rounded, obtuse, or acute; coastal central and southern California (including Channel Islands).</p><!--
+
--><p>latifolia: <i>Grindelia</i> latifolia; stems herbaceous, decumbent to prostrate, leaf blade apices rounded, obtuse, or acute; coastal central and southern California (including Channel Islands).</p><!--
--><p>macrophylla: Grindelia macrophylla; G. integrifolia var. macrophylla; G. stricta var. macrophylla; see stricta.</p><!--
+
--><p>macrophylla: <i>Grindelia</i> macrophylla; <i>G. integrifolia</i> var. macrophylla; G. stricta var. macrophylla; see stricta.</p><!--
--><p>maritima: G. hirsutula var. maritima; G. maritima (Greene) Steyermark; G. rubricaulis var. maritima; stems ascending, 30–80 cm, openly branched, involucres 12–25 mm diam., phyllary apices slightly recurved to nearly straight, cypselae golden or grayish, lengths of pappi ± 1/2 disc corollas; coastal central California. M. A. Lane (1993b) suggested that maritima may have derived from hybridization between hirsutula and platyphylla.</p><!--
+
--><p>maritima: <i>G. hirsutula</i> <i></i>var.<i> maritima</i>; G. maritima (Greene) Steyermark; G. rubricaulis <i></i>var.<i> maritima</i>; stems ascending, 30–80 cm, openly branched, involucres 12–25 mm diam., phyllary apices slightly recurved to nearly straight, cypselae golden or grayish, lengths of pappi ± 1/2 disc corollas; coastal central California. M. A. Lane (1993b) suggested that maritima may have derived from hybridization between hirsutula and platyphylla.</p><!--
--><p>nana: Grindelia nana; stems yellowish to reddish, not resinous, cauline leaf blades usually oblanceolate, lengths usually 5–8 times widths, margins entire or (distally) serrate, involucres mostly 7–10 × 9–15 mm, phyllary apices looped or nearly straight; interior British Columbia, northern California, Idaho, western Montana, northern Nevada, Oregon, Washington.</p><!--
+
--><p>nana: <i>Grindelia</i> nana; stems yellowish to reddish, not resinous, cauline leaf blades usually oblanceolate, lengths usually 5–8 times widths, margins entire or (distally) serrate, involucres mostly 7–10 × 9–15 mm, phyllary apices looped or nearly straight; interior British Columbia, northern California, Idaho, western Montana, northern <i>Nevada</i>, Oregon, Washington.</p><!--
--><p>paludosa: Grindelia paludosa; the nomenclatural type may be a hybrid; stems herbaceous, 50–150 cm, cauline leaf blades coriaceous, little, or not at all, gland-dotted, outer phyllary apices looped, strongly resinous; around Suisun Bay, northern California.</p><!--
+
--><p>paludosa: <i>Grindelia</i> paludosa; the nomenclatural type may be a hybrid; stems herbaceous, 50–150 cm, cauline leaf blades coriaceous, little, or not at all, gland-dotted, outer phyllary apices looped, strongly resinous; around Suisun Bay, northern California.</p><!--
--><p>parviflora: Grindelia camporum var. parviflora; see procera.</p><!--
+
--><p>parviflora: <i>Grindelia</i> camporum <i></i>var.<i> parviflora</i>; see procera.</p><!--
--><p>perennis: Grindelia perennis; perennials, cauline leaf blades oblong to oblanceolate, lengths 6–8 times widths, margins entire or remotely serrulate, phyllary apices strongly resinous, pappi of 2–4 subulate scales; Alberta, British Columbia, Manitoba, Northwest Territories, Ontario, Saskatchewan; Arizona, Colorado, Idaho, Illinois, Michigan, Minnesota, Missouri, Montana, Nebraska, Nevada, New Mexico, New York, North Dakota, Pennsylvania, South Dakota, Utah, Washington, Wyoming.</p><!--
+
--><p>perennis: <i>Grindelia</i> perennis; perennials, cauline leaf blades oblong to oblanceolate, lengths 6–8 times widths, margins entire or remotely serrulate, phyllary apices strongly resinous, pappi of 2–4 subulate scales; Alberta, British Columbia, Manitoba, Northwest Territories, Ontario, Saskatchewan; Arizona, Colorado, Idaho, Illinois, Michigan, Minnesota, Missouri, Montana, Nebraska, <i>Nevada</i>, New Mexico, New York, North Dakota, Pennsylvania, South Dakota, Utah, Washington, Wyoming.</p><!--
--><p>platyphylla: Grindelia latifolia subsp. platyphylla; G. robusta var. platyphylla; G. stricta var. platyphylla; stems herbaceous (caudices woody), greenish, decumbent to prostrate, cauline leaf blades widest proximal to mid points, apices rounded to truncate; dunes and coastal bluffs, central and southern California.</p><!--
+
--><p>platyphylla: <i>Grindelia</i> latifolia subsp. platyphylla; G. robusta var. platyphylla; G. stricta var. platyphylla; stems herbaceous (caudices woody), greenish, decumbent to prostrate, cauline leaf blades widest proximal to mid points, apices rounded to truncate; dunes and coastal bluffs, central and southern California.</p><!--
--><p>procera: Grindelia procera; stems erect, 80–180 cm, strictly branching distally, phyllary apices nearly patent to straight, glabrous, slightly resinous, ray florets 21–45, laminae 8–10 mm; mostly interior central California.</p><!--
+
--><p>procera: <i>Grindelia</i> procera; stems erect, 80–180 cm, strictly branching distally, phyllary apices nearly patent to straight, glabrous, slightly resinous, ray florets 21–45, laminae 8–10 mm; mostly interior central California.</p><!--
--><p>quasiperennis: Grindelia squarrosa var. quasiperennis; see perennis.</p><!--
+
--><p>quasiperennis: <i>Grindelia squarrosa</i> var. quasiperennis; see perennis.</p><!--
--><p>revoluta: Grindelia revoluta; perennials (possibly flowering first year), stems glabrous, cauline leaf blades subcoriaceous, thickened, lengths 2.5–5 times widths, margins entire or remotely dentate to denticulate with short, broad teeth, phyllary apices usually looped, strongly resinous, ray florets 21–37, pappi of 2–4 subulate scales; central and south central Colorado.</p><!--
+
--><p>revoluta: <i>Grindelia</i> revoluta; perennials (possibly flowering first year), stems glabrous, cauline leaf blades subcoriaceous, thickened, lengths 2.5–5 times widths, margins entire or remotely dentate to denticulate with short, broad teeth, phyllary apices usually looped, strongly resinous, ray florets 21–37, pappi of 2–4 subulate scales; central and south central Colorado.</p><!--
--><p>rigida: Grindelia robusta var. rigida; see camporum.</p><!--
+
--><p>rigida: <i>Grindelia</i> robusta <i></i>var.<i> rigida</i>; see camporum.</p><!--
--><p>robusta: Grindelia robusta; fide M. A. Lane (1992b), the nomenclatural type of G. robusta is a hybrid that came from a cross between plants assignable to G. hirsutula in the sense of Lane and plants assignable to G. stricta in the sense of Lane. Within Lane’s interpretation and taxonomic constraints, G. robusta cannot be placed in synonymy of the species name of either “parent.” We believe instead that the type of robusta belongs to the “taxon” that has been called bracteosa.</p><!--
+
--><p>robusta: <i>Grindelia</i> robusta; fide M. A. Lane (1992b), the nomenclatural type of G. robusta is a hybrid that came from a cross between plants assignable to <i>G. hirsutula</i> in the sense of Lane and plants assignable to G. stricta in the sense of Lane. Within Lane’s interpretation and taxonomic constraints, G. robusta cannot be placed in synonymy of the species name of either “parent.” We believe instead that the type of robusta belongs to the “taxon” that has been called bracteosa.</p><!--
--><p>rubricaulis: Grindelia rubricaulis; G. hirsutula subsp. rubricaulis; the type of rubricaulis is very much like the type of hirsutula [see above].</p><!--
+
--><p>rubricaulis: <i>Grindelia</i> rubricaulis; <i>G. hirsutula</i> subsp. rubricaulis; the type of rubricaulis is very much like the type of hirsutula [see above].</p><!--
--><p>stricta: Grindelia stricta; stems usually green to reddish, prostrate to erect, usually ± hirsutulous to arachnose (at least distally), sometimes glabrous, cauline leaf blades usually widest distal to their midpoints, involucres mostly 10–50 mm diam., sometimes subtended by bracteoles, phyllary apices usually slightly recurved to straight, slightly to moderately resinous, ray florets 30–60, laminae 13–25 mm, scales of pappi 0.3+ mm wide at bases; mostly coastal beaches and marshes; British Columbia, Alaska, California, Oregon, Washington.</p><!--
+
--><p>stricta: <i>Grindelia</i> stricta; stems usually green to reddish, prostrate to erect, usually ± hirsutulous to arachnose (at least distally), sometimes glabrous, cauline leaf blades usually widest distal to their midpoints, involucres mostly 10–50 mm diam., sometimes subtended by bracteoles, phyllary apices usually slightly recurved to straight, slightly to moderately resinous, ray florets 30–60, laminae 13–25 mm, scales of pappi 0.3+ mm wide at bases; mostly coastal beaches and marshes; British Columbia, Alaska, California, Oregon, Washington.</p><!--
--><p>venulosa: Grindelia stricta subsp. venulosa; stems procumbent to decumbent, whitish to yellowish, cauline leaf blades fleshy, apices rounded, phyllary apices looped; coastal bluffs and marshes; California, Oregon.</p>
+
--><p>venulosa: <i>Grindelia</i> stricta subsp. venulosa; stems procumbent to decumbent, whitish to yellowish, cauline leaf blades fleshy, apices rounded, phyllary apices looped; coastal bluffs and marshes; California, Oregon.</p>
 
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|publication year=1833
 
|publication year=1833
 
|special status=
 
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/8f726806613d60c220dc4493de13607dd3150896/coarse_grained_fna_xml/V19-20-21/V20_993.xml
 
|tribe=Asteraceae tribe Astereae
 
|tribe=Asteraceae tribe Astereae
 
|genus=Grindelia
 
|genus=Grindelia

Revision as of 15:28, 18 September 2019

Perennials or subshrubs (sometimes flowering first or second year), 8–60(–250+) cm. Stems usually erect, sometimes prostrate, decumbent, or ascending, usually green to stramineous, brown, or reddish, sometimes whitish, arachnose, hirsutulous, puberulous, or villous (sometimes stipitate-glandular as well), or glabrous (then often resinous). Cauline leaf blades usually oblong, oblanceolate, or spatulate (broadest at or beyond their midpoints), sometimes ovate, lanceolate, or linear, (5–)10–80(–120+) mm, lengths 2–8+ times widths, bases clasping or cuneate, margins usually serrate to dentate (teeth apiculate to setose), sometimes entire, apices truncate, rounded, or obtuse to acute, faces hirsutulous, puberulous, or villous and little, if at all, gland-dotted, or glabrous (or scabridulous near margins) and sparsely to densely gland-dotted. Heads usually in open to crowded, corymbiform to paniculiform arrays, seldom borne singly. Involucres usually broadly urceolate to globose, sometimes hemispheric, campanulate, or obconic, (6–)8–15(–20) × (6–)10–25+ mm (sometimes subtended by leaflike bracts). Phyllaries in 4–8+ series, reflexed to spreading or appressed, filiform or linear to ± lance-attenuate, lanceolate, or lance-oblong, apices usually recurved or straight, sometimes (the outer) looped to hooked or patent, terete or filiform to subulate, or acute, usually all glabrous and moderately to strongly resinous, sometimes all or outer villosulous to hirsutulous and little, if at all, resinous, seldom, if ever, stipitate-glandular. Ray florets 0 or (5–)15–60+; laminae (4–)10–25+ mm. Cypselae whitish or stramineous to brownish or grayish, (2–)4–6(–7) mm, apices usually ± knobby, sometimes coronate, rarely smooth, faces usually smooth, striate, or furrowed, rarely rugose; pappi of 2–3(–6) usually contorted or curled, sometimes straight, usually smooth, rarely barbellulate, subulate scales or setiform awns (1–)4–5(–7) mm, usually shorter than, rarely nearly equaling disc corollas. 2n = 12, 24.


Phenology: Flowering year round, mostly (May–)Jul–Sep(–Nov).
Habitat: Disturbed sites, forest openings, hillsides, prairies, roadsides, stream banks, ocean beaches and bluffs, tidal marshes, alkaline, alluvial, clay, or sand soils
Elevation: 0–2800 m

Distribution

V20-993-distribution-map.gif

Alta., B.C., Man., N.W.T., Ont., Que., Sask., Alaska, Ariz., Calif., Colo., Idaho, Ill., Mich., Minn., Mo., Mont., Nebr., Nev., N.Mex., N.Y., N.Dak., Oreg., Pa., S.Dak., Utah, Wash., Wyo., introduced in Mexico (Yucatan).

Discussion

Occurrence of Grindelia hirsutula in Alaska was not verified for this treatment. As circumscribed here, Grindelia hirsutula includes 30 or more reputedly distinct, local, regional, or ecotypic facies that have been named at species or infraspecific rank. Locally, such facies are easily recognized; in a broad view, they intergrade with other facies and are parts of a heterogeneous continuum. Taxonomies that have attempted to recognize the facies as distinct “taxa” have led to almost as many specimens determined as “intermediate” as are assigned to the named “taxa.”

Following, alphabetically by epithet, we have summarized names, diagnostic traits (mostly as given in regional and/or state floras), and distributions for “taxa” treated as distinct in some taxonomies and included within Grindelia hirsutula here:

acutifolia: Grindelia acutifolia; cauline leaf blades firmly membranous, lengths 2.5–5 times widths, phyllary apices looped to hooked, pappi of 2–3, subulate scales; southeastern Colorado, northeastern New Mexico Hybrids between G. acutifolia and G. squarrosa (as G. nuda) have been recorded from the Colorado side of Raton Pass.

altissima: Grindelia nana var. altissima; distal cauline leaves smaller and more scattered than proximal, blades about as wide at bases as at mid points, heads relatively few, heights of involucres less than diams.; mostly inner coastal ranges, northern California, southern Oregon. M. A. Lane (1993b) considered altissima to apply to hybrids between davyi and nana.

angusta: Grindelia inornata var. angusta; cauline leaf blades oblong to oblong-lanceolate, lengths 3+ times widths, faces inconspicuously gland-dotted, ray florets 0, cypselae stramineous to pale brown; central Colorado.

angustifolia: Grindelia stricta var. angustifolia; see humilis.

blakei: Grindelia stricta subsp. blakei; see humilis.

bracteosa: Grindelia bracteosa; G. camporum var. bracteosa; G. robusta var. bracteosa; stems usually whitish and resinous, lengths of cauline leaf blades 3–5 times widths, phyllary apices usually looped, ray florets 0 or 2–27; southern California. M. A. Lane (1993b) suggested that bracteosa may have derived from hybridization between hirsutula and G. squarrosa (var. serrulata). See also, robusta.

camporum: Grindelia camporum; stems usually whitish and resinous, cauline leaf blades ovate to lanceolate, involucres seldom subtended by bracts, phyllary apices recurved to straight, ray florets 32–39; mostly interior north, central California.

columbiana: Grindelia nana [unranked] columbiana; G. columbiana; ray florets 0, otherwise much like nana; Idaho, Oregon, Washington (mostly Columbia River drainage).

davyi: Grindelia camporum var. davyi; G. hirsutula var. davyi; G. robusta var. davyi; phyllary apices recurved to straight, acuminate, pappi usually ± equaling disc corollas; interior northern California. M. A. Lane (1993b) suggested that davyi may have derived from hybridization between camporum and hirsutula.

fastigiata: Grindelia fastigiata; cauline leaf blades conspicuously gland-dotted, phyllary apices thick, coriaceous, ray florets 0, cypselae brown; western Colorado and eastern Utah. Hybrids between fastigiata and G. squarrosa have been recorded from Utah.

hallii: Grindelia hallii; G. hirsutula var. hallii; involucres 8–12 mm diam., phyllary apices slightly recurved and attenuate to nearly straight and deltate, pappi shorter than disc corollas; interior southern California.

hirsutula: Grindelia hirsutula; stems usually green to reddish, ± hirsutulous to arachnose (at least distally), cauline leaf blades usually widest proximal to their midpoints, involucres 12–23 mm diam., often subtended by leaflike bracts, phyllary apices mostly recurved to straight, slightly to moderately resinous, ray florets usually 20–60, laminae 14–20 mm, cypselae usually reddish; mostly coastal ranges of California.

humilis: Grindelia humilis Hooker & Arnott; about the nomenclatural type of G. humilis, M. A. Lane (1992b) stated “...clearly conspecific with G. hirsutula” and she used the name G. stricta var. angustifolia for plants others had treated as G. humilis; as used by others, humilis has been applied to subshrubs 100–150 cm with red-brown stems and phyllary apices recurved to nearly straight from tidal lands around San Francisco and Tomales bays in northern California; similar plants but for apices of phyllaries often looped to hooked from around Humboldt Bay in northern California have been called blakei.

inornata: Grindelia inornata; cauline leaf blades oblanceolate or ovate to oblong, lengths 1.5–3 times widths, faces inconspicuously gland-dotted, ray florets 0, cypselae stramineous to pale brown; central Colorado. Hybrids between inornata and G. subalpina have been recorded from Colorado.

integerrima: Grindelia nana var. integerrima; see integrifolia.

integrifolia: Grindelia nana var. integrifolia Nuttall (not G. integrifolia de Candolle); G. squarrosa var. integrifolia; cauline leaf blades oblanceolate, margins subentire, phyllary apices (at least the outer) looped to hooked, pappi of curled, smooth to barbellulate, subulate scales scarcely longer than cypselae; Idaho, Montana, Nevada, Oregon, Utah, Washington.

latifolia: Grindelia latifolia; stems herbaceous, decumbent to prostrate, leaf blade apices rounded, obtuse, or acute; coastal central and southern California (including Channel Islands).

macrophylla: Grindelia macrophylla; G. integrifolia var. macrophylla; G. stricta var. macrophylla; see stricta.

maritima: G. hirsutula var. maritima; G. maritima (Greene) Steyermark; G. rubricaulis var. maritima; stems ascending, 30–80 cm, openly branched, involucres 12–25 mm diam., phyllary apices slightly recurved to nearly straight, cypselae golden or grayish, lengths of pappi ± 1/2 disc corollas; coastal central California. M. A. Lane (1993b) suggested that maritima may have derived from hybridization between hirsutula and platyphylla.

nana: Grindelia nana; stems yellowish to reddish, not resinous, cauline leaf blades usually oblanceolate, lengths usually 5–8 times widths, margins entire or (distally) serrate, involucres mostly 7–10 × 9–15 mm, phyllary apices looped or nearly straight; interior British Columbia, northern California, Idaho, western Montana, northern Nevada, Oregon, Washington.

paludosa: Grindelia paludosa; the nomenclatural type may be a hybrid; stems herbaceous, 50–150 cm, cauline leaf blades coriaceous, little, or not at all, gland-dotted, outer phyllary apices looped, strongly resinous; around Suisun Bay, northern California.

parviflora: Grindelia camporum var. parviflora; see procera.

perennis: Grindelia perennis; perennials, cauline leaf blades oblong to oblanceolate, lengths 6–8 times widths, margins entire or remotely serrulate, phyllary apices strongly resinous, pappi of 2–4 subulate scales; Alberta, British Columbia, Manitoba, Northwest Territories, Ontario, Saskatchewan; Arizona, Colorado, Idaho, Illinois, Michigan, Minnesota, Missouri, Montana, Nebraska, Nevada, New Mexico, New York, North Dakota, Pennsylvania, South Dakota, Utah, Washington, Wyoming.

platyphylla: Grindelia latifolia subsp. platyphylla; G. robusta var. platyphylla; G. stricta var. platyphylla; stems herbaceous (caudices woody), greenish, decumbent to prostrate, cauline leaf blades widest proximal to mid points, apices rounded to truncate; dunes and coastal bluffs, central and southern California.

procera: Grindelia procera; stems erect, 80–180 cm, strictly branching distally, phyllary apices nearly patent to straight, glabrous, slightly resinous, ray florets 21–45, laminae 8–10 mm; mostly interior central California.

quasiperennis: Grindelia squarrosa var. quasiperennis; see perennis.

revoluta: Grindelia revoluta; perennials (possibly flowering first year), stems glabrous, cauline leaf blades subcoriaceous, thickened, lengths 2.5–5 times widths, margins entire or remotely dentate to denticulate with short, broad teeth, phyllary apices usually looped, strongly resinous, ray florets 21–37, pappi of 2–4 subulate scales; central and south central Colorado.

rigida: Grindelia robusta var. rigida; see camporum.

robusta: Grindelia robusta; fide M. A. Lane (1992b), the nomenclatural type of G. robusta is a hybrid that came from a cross between plants assignable to G. hirsutula in the sense of Lane and plants assignable to G. stricta in the sense of Lane. Within Lane’s interpretation and taxonomic constraints, G. robusta cannot be placed in synonymy of the species name of either “parent.” We believe instead that the type of robusta belongs to the “taxon” that has been called bracteosa.

rubricaulis: Grindelia rubricaulis; G. hirsutula subsp. rubricaulis; the type of rubricaulis is very much like the type of hirsutula [see above].

stricta: Grindelia stricta; stems usually green to reddish, prostrate to erect, usually ± hirsutulous to arachnose (at least distally), sometimes glabrous, cauline leaf blades usually widest distal to their midpoints, involucres mostly 10–50 mm diam., sometimes subtended by bracteoles, phyllary apices usually slightly recurved to straight, slightly to moderately resinous, ray florets 30–60, laminae 13–25 mm, scales of pappi 0.3+ mm wide at bases; mostly coastal beaches and marshes; British Columbia, Alaska, California, Oregon, Washington.

venulosa: Grindelia stricta subsp. venulosa; stems procumbent to decumbent, whitish to yellowish, cauline leaf blades fleshy, apices rounded, phyllary apices looped; coastal bluffs and marshes; California, Oregon.

Selected References

None.

Lower Taxa

None.
... more about "Grindelia hirsutula"
John L. Strother +  and Mark A. Wetter +
Hooker & Arnott +
Alta. +, B.C. +, Man. +, N.W.T. +, Ont. +, Que. +, Sask. +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Ill. +, Mich. +, Minn. +, Mo. +, Mont. +, Nebr. +, Nev. +, N.Mex. +, N.Y. +, N.Dak. +, Oreg. +, Pa. +, S.Dak. +, Utah +, Wash. +, Wyo. +  and introduced in Mexico (Yucatan). +
0–2800 m +
Disturbed sites, forest openings, hillsides, prairies, roadsides, stream banks, ocean beaches and bluffs, tidal marshes, alkaline, alluvial, clay, or sand soils +
Flowering year round, mostly (May–)Jul–Sep(–Nov). +
Bot. Beechey Voy., +
Grindelia acutifolia +, Grindelia camporum +, Grindelia camporum var. bracteosa +, Grindelia camporum var. davyi +, Grindelia camporum var. parviflora +, Grindelia columbiana +, Grindelia fastigiata +, Grindelia hallii +, Grindelia hirsutula var. davyi +, Grindelia hirsutula var. hallii +, Grindelia hirsutula var. maritima +, Grindelia hirsutula subsp. rubricaulis +, Grindelia humilis +, Grindelia inornata +, Grindelia inornata var. angusta +, Grindelia integrifolia var. macrophylla +, Grindelia latifolia +, Grindelia latifolia subsp. platyphylla +, Grindelia macrophylla +, Grindelia maritima +, Grindelia nana +, Grindelia nana var. altissima +, Grindelia nana subsp. columbiana +, Grindelia nana var. integerrima +, Grindelia nana var. integrifolia +, Grindelia paludosa +, Grindelia perennis +, Grindelia procera +, Grindelia revoluta +, Grindelia robusta +, Grindelia robusta var. bracteosa +, Grindelia robusta var. davyi +, Grindelia robusta var. platyphylla +, Grindelia robusta var. rigida +, Grindelia rubricaulis +, Grindelia rubricaulis var. maritima +, Grindelia squarrosa var. integrifolia +, Grindelia squarrosa var. quasiperennis +, Grindelia stricta +, Grindelia stricta var. angustifolia +, Grindelia stricta subsp. blakei +, Grindelia stricta var. macrophylla +, Grindelia stricta var. platyphylla +  and Grindelia stricta subsp. venulosa +
Grindelia hirsutula +
Grindelia +
species +