View source for Geocarpon

{{Treatment/ID
|accepted_name=Geocarpon
|accepted_authority=Mackenzie
|publications={{Treatment/Publication
|title=Torreya
|place=14: 67. 1914
|year=1914
}}
|basionyms=
|synonyms=
|hierarchy=Caryophyllaceae;Caryophyllaceae subfam. Alsinoideae;Geocarpon
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Caryophyllaceae]]</div><div class="higher-taxon"><small>subfamily</small>[[Caryophyllaceae subfam. Alsinoideae]]</div><div class="higher-taxon"><small>genus</small>[[Geocarpon]]</div></div>
|etymology=Greek ge, earth, and carpos, fruit
|volume=Volume 5
|mention_page=page 7
|treatment_page=page 148
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Herbs,</b> annual or winter annual. <b>Taproots</b> slender. <b>Stems</b> erect or spreading-ascending, simple or few-branched basally, terete. <b>Leaves</b> connate basally into sheath, sessile; blade 1-veined, narrowly oblong, not succulent, apex acute. <b>Inflorescences</b> axillary, flowers borne singly at alternate nodes; bracts paired at upper nodes, foliaceous. <b>Pedicels</b>: flowers sessile. <b>Flowers</b>: perianth and androecium perigynous; hypanthium green or reddish, cup-shaped, 10-veined, commisural veins branched ca. at base of sepals; sepals 5, connate proximally ca. 1 mm, green or reddish, 3-veined (lateral pair 1/8 length of midvein), triangular-ovate, margins white, scarious, apex acute; petals absent; nectaries unknown; stamens 5, arising between sepals; filaments distinct; staminodes 5, alternating with stamens, scalelike; styles 3, filiform, ca. 0.3 mm, without stiff hairs proximally; stigmas 3, linear along adaxial surface, minutely papillate (50×). <b>Capsules</b> ovoid, opening by 3 apical, spreading valves; carpophore absent. <b>Seeds</b> ca. 30(–60), yellowish green to brown, translucent, elliptic to ovoid, reniform, slightly laterally compressed, muriculate, marginal wing absent, appendage absent.</span><!--

-->{{Treatment/Body
|distribution=sc United States.
|discussion=<p>Species 1.</p><!--
--><p>When MacKenzie described <i>Geocarpon</i> he placed it in Aizoaceae, thinking that it was closest to <i>Cypselea</i> among the North American representatives of that family. He noted that <i>Geocarpon</i> differs markedly in the absence of stipules and styles and in the valvular rather than circumscissile dehiscence of the fruit. Other authors following this alignment included M. L. Fernald (1950), H. A. Gleason (1952, vol. 2), and P. Wilson (1934). After closer scrutiny of several morphological characteristics, E. J. Palmer and J. A. Steyermark (1950) moved <i>Geocarpon</i> to <i>Caryophyllaceae</i>. Features they noted in suggesting this change included exstipulate leaves, connate sepals, five stamens and, particularly, five staminodes inserted perigynously on the floral tube opposite the calyx lobes, the unilocular ovary with numerous amphitropous ovules on a free-central placenta, and three sessile, recurved stigmas alternating with three bifid tips of the ovary valves. Within <i>Caryophyllaceae</i>, Palmer and Steyermark erected a new monotypic tribe, Geocarpeae, between Alsineae (subtribe Sabulinae) and Sclerantheae. In addition to the morphological evidence for placing <i>Geocarpon</i> in <i>Caryophyllaceae</i>, there are also supporting chemical and anatomical data. The reddish pigment in <i>Geocarpon</i> is an anthocyanin; no betalain-type pigments have been found (A. L. Bogle et al. 1971). Furthermore, H.-D. Behnke (1982) found that the sieve-element plastids of <i>Geocarpon</i> are the type found in <i>Caryophyllaceae</i> (PIIIc´f), not like those found in members of Aizoaceae.</p>
|tables=
|references={{Treatment/Reference
|id=bogle1971a
|text=Bogle, A. L., T. Swain, R. D. Thomas, and E. D. Kohn. 1971. Geocarpon: Aizoaceae or Caryophyllaceae? Taxon 20: 473–477.
}}{{Treatment/Reference
|id=palmer1950a
|text=Palmer, E. J. and J. A. Steyermark. 1950. Notes on Geocarpon minimum Mackenzie. Bull. Torrey Bot. Club 77: 268–273.
}}{{Treatment/Reference
|id=steyermark1959a
|text=Steyermark, J. A., J. W. Voigt, and R. H. Mohlenbrock. 1959. Present biological status of Geocarpon minimum Mackenzie. Bull. Torr. Bot. Club 86: 228–235.
}}
}}<!--

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-->{{#Taxon:
name=Geocarpon
|author=Mark A. Nienaber
|authority=Mackenzie
|rank=genus
|parent rank=subfamily
|synonyms=
|basionyms=
|family=Caryophyllaceae
|illustrator=Barbara Alongi
|illustration copyright=Flora of North America Association
|distribution=sc United States.
|reference=bogle1971a;palmer1950a;steyermark1959a
|publication title=Torreya
|publication year=1914
|special status=
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V5/V5_304.xml
|subfamily=Caryophyllaceae subfam. Alsinoideae
|genus=Geocarpon
}}<!--

-->[[Category:Treatment]][[Category:Caryophyllaceae subfam. Alsinoideae]]