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James D. Montgomery +  and Warren H. Wagner Jr. +
(Villars) H. P. Fuchs +
Polypodium carthusianum +
Spinulose wood fern +, toothed wood fern +  and dryoptère de cartheuser +
Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Ark. +, Conn. +, Del. +, Idaho +, Ill. +, Ind. +, Iowa +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, Mont. +, Nebr. +, N.H. +, N.J. +, N.Y. +, N.C. +, N.Dak. +, Ohio +, Pa. +, R.I. +, S.C. +, Tenn. +, Vt. +, Va. +, Wash. +, W.Va. +, Wis. +  and Eurasia. +
0–1200 m +
Swampy woods, moist wooded slopes, stream banks, and conifer plantations +
Bull. Soc. Bot. France +
Dryopteris austriaca var. spinulosa +, Dryopteris spinulosa +  and Polypodium spinulosum +
Dryopteris carthusiana +
Dryopteris +
species +