View source for Hydrangea

{{Treatment/ID
|accepted_name=Hydrangea
|accepted_authority=Linnaeus
|publications={{Treatment/Publication
|title=Sp. Pl.
|place=1: 397. 1753
|year=1753
}}
|common_names=Graybeard;sevenbark;hortensia
|basionyms=
|synonyms=
|hierarchy=Hydrangeaceae;Hydrangea
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Hydrangeaceae]]</div><div class="higher-taxon"><small>genus</small>[[Hydrangea]]</div></div>
|etymology=Greek hydor, water, and angeion, diminutive of angos, vessel or container, alluding to shape of mature, dehisced capsule
|volume=Volume 12
|mention_page=page 462, 463, 489
|treatment_page=page 486
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Shrubs </b>or trees. <b>Stems</b> erect, ascending, or spreading. <b>Bark</b> exfoliating in grayish, brown, or reddish brown sheets. <b>Branches</b> erect, ascending, or spreading, sometimes arching; twigs with simple or branched trichomes. <b>Leaves</b> deciduous, opposite or 3-whorled; petiole present; blade ovate, elliptic-ovate, elliptic, or suborbiculate, sometimes lobed, herbaceous, margins serrate to serrulate, plane; venation pinnate. <b>Inflorescences</b> terminal, cymose panicles, 100–1000-flowered; peduncle present. <b>Pedicels</b> present. <b>Flowers</b> bisexual or marginal ones often sterile (these with a petaloid, salverform calyx); perianth and androecium epigynous or perigynous; hypanthium campanulate or hemispheric, completely adnate to ovary, weakly or strongly 7–10(–11)-ribbed in fruit; sepals persistent, 5, spreading or reflexed, deltate to shallowly triangular, usually glabrous, rarely abaxially sparsely hairy; petals 5, valvate, spreading or reflexed, white to yellowish white, ovate-lanceolate, elliptic, oblong, spatulate, or narrowly ovate to ovate, base sessile, surfaces glabrous; stamens 10; filaments distinct, terete or flattened proximally, gradually or abruptly tapered from base to apex, apex not 2-lobed; anthers depressed-ovoid or transversely oblong; pistil 2–4-carpellate, ovary completely inferior or nearly so, or 1/2 inferior, 2–4-locular; placentation axile proximally, parietal distally; styles persistent, 2–4, distinct or connate to middle or distally. <b>Capsules</b> hemispheric, suburceolate, or oblong-ovoid, coriaceous, dehiscence interstylar, creating elliptic to circular pore at base of styles. <b>Seeds</b> 10–40 per locule, light brown to dark brown, fusiform or ellipsoid. <b>x</b> = 18.</span><!--

-->{{Treatment/Body
|distribution=United States;Mexico;Central America;w South America;e Asia.
|discussion=<p>Species ca. 29 (5 in the flora).</p><!--
--><p><i>Hydrangea</i> enjoys considerable esteem as an ornamental shrub, especially for its prominent sterile flowers. North American species have been cultivated in Europe since before the mid 1700s (W. L. Stern 1978). Besides the species treated here, popular ornamentals in North America are H. anomala D. Don, H. aspera D. Don, H. heteromalla D. Don, H. involucrata Siebold, H. macrophylla (Thunberg) Seringe, and H. serrata (Thunberg) Seringe. Among these, H. macrophylla may be the most widely grown; M. A. Dirr (2004) listed nearly 170 cultivars of this species. Surprisingly, it has not escaped from cultivation.</p><!--
--><p>Tubercles, comprising crystals of calcium carbonate, often are visible on leaf trichomes (G. W. Burkett 1932). They are observed most easily at magnifications greater than 30\x, and some taxonomic utility has been ascribed to their presence and abundance.</p><!--
--><p>Toxic and medicinal properties are attributed to some native and cultivated species of <i>Hydrangea</i> (J. M. Kingsbury 1964; W. L. Stern 1978; D. E. Moerman 1998; G. E. Burrows and R. J. Tyrl 2001). These possibly are related to various alkaloids present in roots and leaves of some species.</p><!--
--><p>Y. De Smet et al. (2015) found <i>Hydrangea</i> to be polyphyletic and promoted adoption of a broader, monophyletic concept of <i>Hydrangea</i> that includes all eight genera in tribe Hydrangeeae. Both North American genera in the tribe, <i>Decumaria</i> and <i>Hydrangea</i>, are circumscribed here in their traditional senses.</p>
|tables=
|references={{Treatment/Reference
|id=dirr2004a
|text=Dirr, M. A. 2004. Hydrangeas for American Gardens. Portland.
}}{{Treatment/Reference
|id=mcclintock1957a
|text=McClintock, E. 1957. A monograph of the genus Hydrangea. Proc. Calif. Acad. Sci., ser. 4, 29: 147–256.
}}{{Treatment/Reference
|id=pilatowski1980a
|text=Pilatowski, R. E. 1980. A Taxonomic Study of the Hydrangea arborescens Complex. M.S. thesis. North Carolina State University.
}}{{Treatment/Reference
|id=pilatowski1982a
|text=Pilatowski, R. E. 1982. A taxonomic study of the Hydrangea arborescens complex. Castanea 47: 84–98.
}}{{Treatment/Reference
|id=st1921a
|text=St. John, H. 1921. A critical revision of Hydrangea arborescens. Rhodora 23: 203–208.
}}{{Treatment/Reference
|id=stern1978a
|text=Stern, W. L. 1978. Comparative anatomy and systematics of woody Saxifragaceae. Hydrangea. Bot. J. Linn. Soc. 76: 83–113.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Leaf blades (3–)5(–7)-lobed, margins coarsely serrate; inflorescences ovoid to conic.
|[[Hydrangea quercifolia|Hydrangea quercifolia]]
|-id=key-0-1
|1
|Leaf blades unlobed, margins dentate, serrate, or serrulate; inflorescences dome-shaped to hemispheric or conic to ovoid.
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Ovaries 1/2 inferior; inflorescences usually conic, sometimes ovoid.
|[[Hydrangea paniculata|Hydrangea paniculata]]
|-id=key-0-2
|2
|Ovaries completely inferior or nearly so; inflorescences dome-shaped to hemispheric.
|[[#key-0-3| > 3]]
|-id=key-0-3
|3
|Leaf blade abaxial surfaces white or grayish, densely tomentose, trichomes at 40× either smooth, 1–3 mm, or sparsely tuberculate, 0.3–1 mm.
|[[Hydrangea radiata|Hydrangea radiata]]
|-id=key-0-3
|3
|Leaf blade abaxial surfaces green, glabrous or glabrate, or grayish, uniformly velutinous, pilose, or tomentose, trichomes at 40× conspicuously tuberculate, 0.3–1 mm.
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Leaf blade abaxial surfaces green, glabrous or glabrate, or sparsely hirsute along midveins and sometimes along lateral veins.
|[[Hydrangea arborescens|Hydrangea arborescens]]
|-id=key-0-4
|4
|Leaf blade abaxial surfaces grayish, uniformly velutinous, pilose, or tomentose.
|[[Hydrangea cinerea|Hydrangea cinerea]]
|}
</div></div><!--

-->{{#Taxon:
name=Hydrangea
|author=Craig C. Freeman
|authority=Linnaeus
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Hydrangeaceae
|distribution=United States;Mexico;Central America;w South America;e Asia.
|reference=dirr2004a;mcclintock1957a;pilatowski1980a;pilatowski1982a;st1921a;stern1978a
|publication title=Sp. Pl.
|publication year=1753
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/8f726806613d60c220dc4493de13607dd3150896/coarse_grained_fna_xml/V12/V12_115.xml
|genus=Hydrangea
}}<!--

-->[[Category:Treatment]][[Category:Hydrangeaceae]]