View source for Achlys

{{Treatment/ID
|accepted_name=Achlys
|accepted_authority=de Candolle
|publications={{Treatment/Publication
|title=Syst. Nat.
|place=2: 35. 1821
|year=1821
}}
|basionyms=
|synonyms=
|hierarchy=Berberidaceae;Achlys
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Berberidaceae]]</div><div class="higher-taxon"><small>genus</small>[[Achlys]]</div></div>
|etymology=Greek Achlus, a god of night
|volume=Volume 3
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Herbs,</b> perennial, deciduous, 2.5-5 dm, glabrous. <b>Rhizomes</b> extensive, branching, producing 1-few foliage leaves or flowering shoots per year. <b>Aerial</b> stems absent. <b>Leaves</b> basal, alternate, 3-foliolate; petiole long, slender. <b>Leaf</b> blade orbiculate in gross outline, 20-40 cm; leaflet blades fan-shaped, entire or lobed, lateral leaflet blades strongly asymmetric, margins entire to coarsely sinuate; venation palmate. <b>Inflorescences</b> terminal, dense scapose-pedunculate spikes of inconspicuous flowers. <b>Flowers</b> 3-merous, white to cream, 6 mm or less; bracteoles absent; sepals absent; petals absent; stamens 8-10(-13); anthers dehiscing by 2 apically hinged flaps; pollen exine striate; ovaries asymmetrically ellipsoid; placentation marginal, placenta developed only near base of ovary. <b>Fruits</b> follicles with transverse dehiscence, purplish red or brown, curved, furrowed. <b>Seeds</b> 1, brown; aril absent. <b>x</b> = 6.</span><!--

-->{{Treatment/Body
|distribution=North America;Asia (Japan).
|discussion=<p>Species 3 (2 in the flora).</p><!--
--><p>Achlys is of particular interest because of its amphi-Pacific disjunction. Despite the 8000 km or more disjunction, the taxa are remarkably similar in morphology, ecology, and karyology. Japanese populations are diploid; American populations are diploid and tetraploid.</p><!--
--><p>Two species are recognized in this treatment; some researchers prefer to treat them as varieties. In the Californian portion of the range, some field botanists believe the two taxa are sufficiently morphologically distinct to be called species; farther north these differences are reportedly less apparent.</p>
|tables=
|references={{Treatment/Reference
|id=fukuda1967a
|text=Fukuda, I. 1967. The biosystematics of Achlys. Taxon 16: 308-316.
}}{{Treatment/Reference
|id=fukuda1970a
|text=Fukuda, I. and H. G. Baker. 1970. Achlys californica --(Berberidaceae)--a new species. Taxon 19: 341-344.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Central leaflet blade 4–11 cm, distal margins (1–)3–4(–8)-lobed; stamens 3–4 mm; follicles red-purple.
|[[Achlys triphylla|Achlys triphylla]]
|-id=key-0-1
|1
|Central leaflet blade ca. 7–16 cm, distal margins (3–)6–9(–12)-lobed; stamens 3.8–6 mm; follicles brown.
|[[Achlys californica|Achlys californica]]
|}
</div></div><!--

-->{{#Taxon:
name=Achlys
|author=R. David Whetstone; T.A. Atkinson; Daniel D. Spaulding
|authority=de Candolle
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Berberidaceae
|distribution=North America;Asia (Japan).
|reference=fukuda1967a;fukuda1970a
|publication title=Syst. Nat.
|publication year=1821
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V3/V3_251.xml
|genus=Achlys
}}<!--

-->[[Category:Treatment]][[Category:Berberidaceae]]