View source for Hedwigia

{{Treatment/ID
|accepted_name=Hedwigia
|accepted_authority=P. Beauvois
|publications={{Treatment/Publication
|title=Mag. Encycl.
|place=5: 304. 1804
|year=1804
}}
|basionyms=
|synonyms=
|hierarchy=Hedwigiaceae;Hedwigia
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Hedwigiaceae]]</div><div class="higher-taxon"><small>genus</small>[[Hedwigia]]</div></div>
|etymology=For Johann Hedwig, 1730 – 1799, German bryologist and physician
|volume=Volume 28
|mention_page=page 85, 88, 90, 642
|treatment_page=page 84
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>large, sordid yellow-green, rarely bright green, pale buff to red-brown with age, sometimes hoary distally. <b>Stems</b> with stoloniform-flagelliform branches absent. <b>Leaves</b> erect to subsecund and imbricate, not undulate-plicate, occasionally somewhat striolate when dry, 1.5–3 mm; margins recurved or sometimes plane in proximal 1/2–2/3, plane, erect, or incurved in acumen, in larger leaves closely and irregularly dentate to spinulose-dentate in apex; apex erect to patent when dry, wide-spreading and recurved or reflexed when moist, acute, acuminate, or subpiliferous, concolorous when muticous, or hyaline-white with pellucid tip, canaliculate-furrowed to tubular, papillose proximally, spinose-serrate medially, smooth apically; alar cells oblate, quadrate, or short-rectangular, smooth to simple-papillose, walls even to thick and porose, region concolorous or color similar to mid basal region, in several rows along margins; mid basal laminal cells long-rectangular, papillae 4–7, in 1 row, walls strongly thick, moderately to strongly porose, region strongly yellow- to red-orange across insertion or to 1/4–1/3(–1/2) leaf length; medial cells 1- or multipapillose on both surfaces, papillae simple or multifid, walls thick, moderately to coarsely and irregularly porose-sinuate; distal cells chlorophyllose and similar to medial cells, or echlorophyllose and differentiated, prorulose at distal cell ends, papillose; apical cells pellucid, less papillose. <b>Sexual</b> condition autoicous; perichaetial leaves to 4 mm, margins entire or ciliate distally. <b>Vaginula</b> with paraphyses few to many, smooth, occasionally papillose, often extending onto developing calyptra. <b>Seta</b> reddish brown, 0.5–0.9 mm, stout, hidden by perichaetial leaves. <b>Capsule</b> deeply immersed, brown proximally, shiny red-brown at mouth, turbinate-urceolate when dry, subglobose, short-ovoid, or obovoid when moist, 1 mm, longitudinally wrinkled to sharply sulcate when dry, smooth except for neck, mouth wide; stomata cryptoporic; operculum planoconvex, sometimes subumbonate-apiculate. <b>Calyptra</b> conic-mitrate, 0.5–0.9 mm, covering apex of capsule, pilose or naked. <b>Spores</b> 19–30 µm, finely to somewhat coarsely vermiculate-papillose.</span><!--

-->{{Treatment/Body
|distribution=Nearly worldwide.
|discussion=<p>Species 6 (3 in the flora).</p><!--
--><p><i>Hedwigia</i> in North America was until recently considered as a single species, with numerous varieties and forms (G. N. Jones 1933). Although a hair-point is often ascribed to species of <i>Hedwigia</i>, the acumen may be elongated but there is never a hairlike process as in <i>Pseudobraunia</i>. Long, multicellular cilia on leaf margins are rare among bryophytes (B. Goffinet et al. 2008), but are present on the perichaetial leaves in <i>H. ciliata</i> and <i>H. stellata</i>.</p><!--
--><p>The type of <i>Hedwigia</i> integrifolia P. Beauvois was collected in North America. The name was used for a widespread species now considered <i>Braunia</i> imberbis (Smith) N. Dalton & D. G. Long. The type of <i>H. integrifolia</i> was examined by N. J. Dalton et al. (2012) and found unrelated to B. imberbis. <i>Hedwigia</i> integrifolia was synonymized with Hedwigidium imberbe (Smith) Bruch & Schimper by W. J. Hooker and T. Taylor (1818). Presently the application of the name <i>H. integrifolia</i> is awaiting further study, although G. N. Jones (1933) considered <i>H. integrifolia</i> a synonym of <i>H. ciliata</i> (B. H. Allen 2010).</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Medial and distal laminal cells with papillae (1-)2-4 per cell, simple, sessile, or low-stalked, variously branched; perichaetial leaf margins ciliate distally; apical laminal cell obtuse to truncate, multipapillose-coronate, short-rhomboidal, (40-)50-75(-80) µm.
|[[Hedwigia ciliata|Hedwigia ciliata]]
|-id=key-0-1
|1
|Medial and distal laminal cells with papillae usually 1 per cell, stalked, strongly, irregularly branched; perichaetial leaf margins ciliate or entire distally; apical laminal cell sharply pointed, smooth or papillose, long-linear, (80-)120-175(-200) µm
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Leaf apices wide-spreading to squarrose when dry; laminal cells in hyaline area with some papillae large, spinose, among shorter papillae; papillae on abaxial leaf surface with branches of unequal lengths; perichaetial leaf margins long-ciliate distally.
|[[Hedwigia stellata|Hedwigia stellata]]
|-id=key-0-2
|2
|Leaf apices erect when dry; laminal cells in hyaline area with papillae low, simple, in lines; papillae on abaxial leaf surface with branches usually of equal lengths; perichaetial leaf margins entire.
|[[Hedwigia detonsa|Hedwigia detonsa]]
|}
</div></div><!--

-->{{#Taxon:
name=Hedwigia
|authority=P. Beauvois
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Hedwigiaceae
|distribution=Nearly worldwide.
|reference=None
|publication title=Mag. Encycl.
|publication year=1804
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-data-curation.git/src/bb6b7e3a7de7d3b7888a1ad48c7fd8f5c722d8d6/coarse_grained_fna_xml/V28/V28_132.xml
|genus=Hedwigia
}}<!--

-->[[Category:Treatment]][[Category:Hedwigiaceae]]