View source for Tolmiea ← Tolmiea You do not have permission to edit this page, for the following reason: The action you have requested is limited to users in the group: Users. You can view and copy the source of this page. {{Treatment/ID |accepted_name=Tolmiea |accepted_authority=Torrey & A. Gray |publications={{Treatment/Publication |title=Fl. N. Amer. |place=1: 582. 1840, name conserved , }} |common_names=Pig-a-back plant |basionyms= |synonyms= |hierarchy=Saxifragaceae;Tolmiea |hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Saxifragaceae]]</div><div class="higher-taxon"><small>genus</small>[[Tolmiea]]</div></div> |etymology=For William F. Tolmie, 1812–1886, surgeon for Hudson’s Bay Company at Fort Vancouver |volume=Volume 8 |mention_page=page 43, 44, 45, 46, 108, 109 |treatment_page=page 107 }}<!-- --><span class="statement" id="st-undefined" data-properties=""><b>Herbs,</b> rhizomatous, not stoloniferous; rhizomes sometimes branched, scaly; caudex not cormlike, with persistent leaf bases. <b>Flowering</b> stems erect, leafy, 2.5–70 cm, sparsely to moderately, short to long stipitate-glandular. <b>Leaves</b> in basal rosette and cauline; cauline leaves reduced distally, vegetative shoots sometimes from axillary buds of cauline leaves; stipules present; petiole stipitate-glandular, (adventitious buds usually produced at apices of petioles of rosette and cauline leaves, sometimes forming plantlets); blade ovate, shallowly 5–9-lobed, base cordate, ultimate margins strongly to obscurely, irregularly serrate to nearly dentate, each tooth ending in glandular hair, apex acute (rarely obtuse), surfaces subglabrous to stipitate-glandular; venation palmate. <b>Inflorescences</b> racemes, from axillary buds in rosette, 10–150-flowered, bracteate. (Pedicels with subtending bracts.) Flowers bilaterally symmetric; hypanthium adnate to ovary only at base for less than 1 mm (ovary appearing superior), free from ovary for 4–5 mm, greenish, ± split to base, (cylindric-funnelform); sepals (persistent), 5, green with variable development of red-purple stripes, (unequal, slightly gibbous at base, stipitate-glandular, 3 dorsal sepals ovate- to elliptic-triangular, apex obtuse to rounded-mucronate, 2 ventral-lateral sepals narrowly ovate-triangular to oblong-triangular, apex acute to acuminate); petals 4, (recurved), greenish to brown-purple, (± linear); nectariferous tissue proximal to stamens; stamens 3, (inserted at apex of hypanthium opposite dorsal sepals, slightly exserted); filaments filiform; ovary superior, 1-locular, (oblong, turbinate, apex 2-cleft, stipitate-glandular), carpels connate proximally; placentation parietal; styles 2; stigmas 2. <b>Capsules</b> 2-beaked, (ovoid, turbinate). <b>Seeds</b> brown to nearly black, subglobose, distinctly muricate. <b>x</b> = 7.</span><!-- -->{{Treatment/Body |distribution=nw North America. |discussion=<p>Species 2 (2 in the flora).</p><!-- --><p>Although morphologically similar, as reviewed by D. E. Soltis et al. (2007), diploid and tetraploid Tolmiea each meets the criteria for all of the commonly used species concepts: they are reproductively isolated, so they are biological species; they are distinct evolutionary lineages, so the two cytotypes meet the expectations of the evolutionary species concept; each is monophyletic, so both can be considered phylogenetic (apomorphic) species; they are diagnosable on the basis of chromosome number and molecular characters, thus fitting the phylogenetic/diagnosability species concept; and morphological characters distinguish the two cytotypes (although these may be considered cryptic by some), hence they are taxonomic species.</p> |tables= |references={{Treatment/Reference |id=soltis1984a |text=Soltis, D. E. 1984b. Autopolyploidy in Tolmiea menziesii (Saxifragaceae). Amer. J. Bot. 71: 1171–1174. }}{{Treatment/Reference |id=soltis1988a |text=Soltis, D. E. and P. S. Soltis. 1988. Electrophoretic evidence for tetrasomic inheritance in Tolmiea menziesii (Saxifragaceae). Heredity 60: 375–382. }}{{Treatment/Reference |id=soltis1989a |text=Soltis, D. E. and P. S. Soltis. 1989. Genetic consequences of autopolyploidy in Tolmiea (Saxifragaceae). Evolution 43: 586–594. }} }}<!-- --><div class="treatment-key"> ==Key== <div class="treatment-key-group"> {| class="wikitable fna-keytable" |-id=key-0-1 |1 |Cauline leaves not or only slightly longer than wide, i.e., length-width quotient usually 0.6-1.1 (average 0.9, excluding length of basal lobes); plantlets frequently formed. |[[Tolmiea menziesii|Tolmiea menziesii]] |-id=key-0-1 |1 |Cauline leaves distinctly longer than wide, i.e., length-width quotient 0.8-1.4 (average 1, excluding length of basal lobes); plantlets on leaves sporadically produced. |[[Tolmiea diplomenziesii|Tolmiea diplomenziesii]] |} </div></div><!-- -->{{#Taxon: name=Tolmiea |author=Douglas E. Soltis;Walter S. Judd;Pamela S. Soltis;Patrick E. Elvander† |authority=Torrey & A. Gray |rank=genus |parent rank=family |synonyms= |basionyms= |family=Saxifragaceae |distribution=nw North America. |reference=soltis1984a;soltis1988a;soltis1989a |publication title=Fl. N. Amer. |publication year= |special status= |source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V8/V8_212.xml |genus=Tolmiea }}<!-- -->[[Category:Treatment]][[Category:Saxifragaceae]] Templates used on this page: Template:Saxifragaceae (view source) Template:Treatment/AuthorLink (view source) Template:Treatment/Body (view source) Template:Treatment/Body/Maps (view source) Template:Treatment/ID (view source) Template:Treatment/Publication (view source) Template:Treatment/Reference (view source) Return to Tolmiea.