View source for Horkelia sect. Horkelia

{{Treatment/ID
|accepted_name=Horkelia sect. Horkelia
|accepted_authority=
|publications=
|basionyms=
|synonyms={{Treatment/ID/Synonym
|name=Californicae
|authority=Rydberg
|rank=unranked
}} {{Treatment/ID/Synonym
|name=Horkelia sect. Californicae
|authority=(Rydberg) O. Stevens
|rank=section
}} {{Treatment/ID/Synonym
|name=Cuneatae
|authority=Rydberg
|rank=unranked
}} {{Treatment/ID/Synonym
|name=Horkelia sect. Cuneatae
|authority=(Rydberg) O. Stevens
|rank=section
}} {{Treatment/ID/Synonym
|name=Tenuilobae
|authority=Rydberg
|rank=unranked
}} {{Treatment/ID/Synonym
|name=Horkelia sect. Tenuilobae
|authority=
|rank=section
}}
|hierarchy=Rosaceae;Rosaceae subfam. Rosoideae;Rosaceae tribe Potentilleae;Horkelia;Horkelia sect. Horkelia
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Rosaceae]]</div><div class="higher-taxon"><small>subfamily</small>[[Rosaceae subfam. Rosoideae]]</div><div class="higher-taxon"><small>tribe</small>[[Rosaceae tribe Potentilleae]]</div><div class="higher-taxon"><small>genus</small>[[Horkelia]]</div><div class="higher-taxon"><small>section</small>[[Horkelia sect. Horkelia]]</div></div>
|volume=Volume 9
|mention_page=page 248, 259
|treatment_page=page 250
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>forming tufts or mats, green to grayish, obscurely (and minutely) glandular, resinously aromatic, often strongly so. <b>Stems</b> decumbent or ascending to erect, (0.5–)1–10(–12) dm. <b>Basal</b> leaves usually planar, sometimes ± cylindric; stipules usually entire, sometimes basally lobed; leaflets (1–)3–16(–20) per side, separate to overlapping, divided ± 1/6–3/4+ to midrib into 3–30(–60) teeth or lobes not restricted to apex. <b>Inflorescences</b> open to congested, flowers arranged individually, in usually non-capitate glomerules, and/or in corymbiform clusters. <b>Pedicels</b> remaining straight, 1–30(–40) mm. <b>Flowers</b>: epicalyx bractlets narrowly elliptic-lanceolate to broadly ovate, 0.5–3 mm wide, usually entire, sometimes toothed; hypanthium interior pilose or glabrous; sepals acute; petals white, oblong-oblanceolate to round, apex obtuse to truncate to emarginate; filaments white, glabrous, anthers longer than wide; carpels 10–200(–220). <b>Achenes</b> 0.8–2 mm, usually smooth or slightly rugose, sometimes merely roughened.</span><!--

-->{{Treatment/Body
|distribution=Calif.;nw Mexico.
|discussion=<p>Species 9 (9 in the flora).</p><!--
--><p><i>Section</i> <i>Horkelia</i> encompasses the species that are most commonly encountered in heavily populated areas of California. Plants are notably glandular-viscid (unless obscured by dense vestiture) and have a distinctive resinous odor.</p><!--
--><p>Previous revisions (for example, P. A. Rydberg 1908c; D. D. Keck 1938) have placed <i>Horkelia</i> frondosa (here treated as <i>H. californica </i>var.<i> frondosa</i>) at the beginning, implying that this is the least derived expression within the genus. Such an assumption is based on its gross resemblance to sympatric members of <i>Drymocallis</i>; molecular evidence (T. Eriksson et al. 1998; M. Lundberg et al. 2009; C. Dobeš and J. Paule 2010) confirms that this similarity is superficial. If, as speculated above, species composing sect. Hispidulae are relicts of the original radiation, then <i>H. californica </i>var.<i> frondosa</i> is actually one of the more derived members of the genus. Species within sect. <i>Horkelia</i> have been ordered here according to that interpretation.</p>
|tables=
|references={{Treatment/Reference
|id=ertter1993a
|text=Ertter, B. 1993c. A re-evaluation of the Horkelia bolanderi (Rosaceae) complex, with the new species Horkelia yadonii. Syst. Bot. 18: 137–144.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Leaflets 1–3 per side, sparsely short-pilose to glabrate, (15–)20–30-toothed 1/6–1/4 to midrib; petals obovate to round, (3–)4–7 mm wide.
|[[Horkelia truncata|Horkelia truncata]]
|-id=key-0-1
|1
|Leaflets (3–)5–16(–20) per side, hairy, 3–15(–30)-toothed or -lobed (collectively 10–60-toothed in H. californica) 1/6–3/4+ to midrib; petals narrowly obovate or oblanceolate to elliptic or oblong, 1–3(–4) mm wide
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Hypanthia 3–5.5 mm, 1/2 to nearly as deep as wide; epicalyx bractlets 4–6(–8) mm, ± equal to sepals, entire or toothed.
|[[Horkelia californica|Horkelia californica]]
|-id=key-0-2
|2
|Hypanthia 1–2(–3) mm, less than 1/2 as deep as wide (except H. clevelandii); epicalyx bractlets 1–4(–5) mm, shorter than sepals, entire
|[[#key-0-3| > 3]]
|-id=key-0-3
|3
|Leaflets ± elliptic to obovate, divided less than 1/3 to midrib into (5–)10–15 teeth; epicalyx bractlets 1.5–3 mm wide.
|[[Horkelia cuneata|Horkelia cuneata]]
|-id=key-0-3
|3
|Leaflets cuneate or flabellate to ovate or nearly round, divided (1/5–)1/3–3/4+ to midrib into 3–10(–30 in H. yadonii) teeth or lobes; epicalyx bractlets 0.5–2(–2.5) mm wide
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Stems ± decumbent to ascending, 1–3.5(–4.5) dm; leaflets 5–10(–12) per side, cuneate, ± overlapping; inflorescences usually congested; plants matted; coastal.
|[[Horkelia marinensis|Horkelia marinensis]]
|-id=key-0-4
|4
|Stems ascending to erect, (0.5–)1–7 dm; leaflets 6–16(–20) per side, cuneate to flabellate or nearly round, separate to ± overlapping; inflorescences open to ± congested; plants tufted to matted; coastal or interior
|[[#key-0-5| > 5]]
|-id=key-0-5
|5
|Epicalyx bractlets 1–2(–2.5) mm wide, lanceolate to ovate; anthers 0.8–1.2 mm; hypanthia 3–6.5 mm diam.
|[[Horkelia yadonii|Horkelia yadonii]]
|-id=key-0-5
|5
|Epicalyx bractlets 0.5–1(–1.5) mm wide, narrowly elliptic to broadly lanceolate; anthers 0.4–1 mm; hypanthia 2–4.5(–6) mm diam
|[[#key-0-6| > 6]]
|-id=key-0-6
|6
|Stems: hairs ascending to appressed; plants usually grayish to grayish green; hypanthium interior pilose
|[[#key-0-7| > 7]]
|-id=key-0-6
|6
|Stems: hairs ± spreading; plants green to grayish; hypanthium interior glabrous or pilose
|[[#key-0-8| > 8]]
|-id=key-0-7
|7
|Carpels 20–50(–120); styles 2–4 mm; basal leaves (4–)8–20(–30) cm; Transverse Ranges, s California.
|[[Horkelia rydbergii|Horkelia rydbergii]]
|-id=key-0-7
|7
|Carpels 10–20(–27); styles (1–)1.5–2 mm; basal leaves 3–8(–9) cm; Coast Ranges, n California.
|[[Horkelia bolanderi|Horkelia bolanderi]]
|-id=key-0-8
|8
|Leaflets divided 1/2–3/4+ to midrib into linear to narrowly oblanceolate or narrowly elliptic lobes; hypanthium interior pilose; n Coast Ranges, California.
|[[Horkelia tenuiloba|Horkelia tenuiloba]]
|-id=key-0-8
|8
|Leaflets divided 1/3 to midrib into acute to obtuse teeth; hypanthium interior glabrous; Peninsular Ranges, s California.
|[[Horkelia clevelandii|Horkelia clevelandii]]
|}
</div></div><!--

-->{{#Taxon:
name=Horkelia sect. Horkelia
|authority=
|rank=section
|parent rank=genus
|synonyms=Californicae;Horkelia sect. Californicae;Cuneatae;Horkelia sect. Cuneatae;Tenuilobae;Horkelia sect. Tenuilobae
|basionyms=
|family=Rosaceae
|distribution=Calif.;nw Mexico.
|reference=ertter1993a
|publication title=
|publication year=
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/f50eec43f223ca0e34566be0b046453a0960e173/coarse_grained_fna_xml/V9/V9_390.xml
|subfamily=Rosaceae subfam. Rosoideae
|tribe=Rosaceae tribe Potentilleae
|genus=Horkelia
|section=Horkelia sect. Horkelia
}}<!--

-->[[Category:Treatment]][[Category:Horkelia]]