View source for Hydrastis

{{Treatment/ID
|accepted_name=Hydrastis
|accepted_authority=J. Ellis in C. Linnaeus
|publications={{Treatment/Publication
|title=Syst. Nat. ed.
|place=10, 2: 1088. 1759
|year=1759
}}
|common_names=Goldenseal;orangeroot;yellow-puccoon;sceau d'or
|basionyms=
|synonyms=
|hierarchy=Ranunculaceae;Hydrastis
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Ranunculaceae]]</div><div class="higher-taxon"><small>genus</small>[[Hydrastis]]</div></div>
|etymology=referring to superficial resemblance to some species of Hydrophyllum
|volume=Volume 3
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Herbs,</b> perennial, from creeping, yellow, thick rhizomes. <b>Leaves</b> basal and cauline, simple, petiolate; cauline leaves alternate. <b>Leaf</b> blade palmately 3-9-lobed [variously divided], broadly cordate-orbiculate, margins serrate. <b>Inflorescences</b> terminal, solitary flowers; bracts absent. <b>Flowers</b> bisexual, radially symmetric; sepals not persistent in fruit, 3, greenish white to creamy white, plane, ovate, oval, or elliptic, 3.5-7 mm; petals absent; stamens 50-75; filaments abruptly narrowed near apex; staminodes absent between stamens and pistils; pistils 5-15, simple; ovules 2 per ovary; style short. <b>Fruits</b> berries, aggregate, sessile, spheric, sides not veined; beak terminal, ± straight, 0.6-1 mm. <b>Seeds</b> black, ellipsoid, smooth, lustrous. <b>x</b> = 13.</span><!--

-->{{Treatment/Body
|distribution=e North America;Asia (Japan).
|discussion=<p>Species 2 (1 in the flora).</p><!--
--><p>Many classifications have included <i>Hydrastis</i> within Ranunculaceae (A. Cronquist 1981; W. C. Gregory 1941; O. F. Langlet 1932; M. Tamura 1963, 1968, 1993). Recent workers (H. Tobe and R. C. Keating 1985; A. L. Takhtajan 1987), however, have assigned <i>Hydrastis</i> to its own family, intermediate between Ranunculaceae and Berberidaceae. Phylogenetic studies by S. B. Hoot (1991) confirmed the isolated position of <i>Hydrastis</i> and suggested that characteristics such as the following justify its exclusion from Ranunculaceae: undifferentiated mesophyll, xylem straight (instead of V-shaped) in cross section, scalariform perforations in the vessels, micropyle defined by two integuments, pollen with a distinct striate-reticulate tectum, and a base chromosome number of x = 13 (as opposed to 6, 7, 8, or 9).</p><!--
--><p>C. S. Keener (1993) challenged these conclusions, stating that features such as scalariform perforations in vessels, striate-reticulate pollen, and the micropyle defined by two integuments are found in other genera within Ranunculaceae. Only straight xylem in cross section and the base chromosome number are distinctive. Whether these features warrant segregation of <i>Hydrastis</i> into its own family is debatable. Decisions involving the circumscription of this genus await a molecular study involving Berberidaceae, Ranunculaceae, and related families.</p>
|tables=
|references={{Treatment/Reference
|id=eichenberger1976a
|text=Eichenberger, M. D. and G. R. Parker. 1976. Goldenseal (Hydrastis canadensis L.) distribution, phenology and biomass in an oak-hickory forest. Ohio J. Sci. 76: 204-210.
}}{{Treatment/Reference
|id=keener1993a
|text=Keener, C. S. 1993. A review of the classification of the genus Hydrastis (Ranunculaceae). Aliso 13: 551-558.
}}{{Treatment/Reference
|id=massey1983a
|text=Massey, J. R., D. K. S. Otte, T. A. Atkinson, and R. D. Whetstone. 1983. An Atlas and Illustrated Guide to Threatened and Endangered Vascular Plants of the Mountains of North Carolina and Virginia. Washington. [U.S.D.A. Forest Serv., Gen. Techn. Rep. SE-20.]
}}{{Treatment/Reference
|id=tobe1985a
|text=Tobe, H. and R. C. Keating. 1985. The morphology and anatomy of Hydrastis (Ranunculales): Systematic reevaluation of the genus. Bot. Mag. (Tokyo) 98: 291-316.
}}
}}<!--

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-->{{#Taxon:
name=Hydrastis
|author=
|authority=J. Ellis in C. Linnaeus
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Ranunculaceae
|distribution=e North America;Asia (Japan).
|reference=eichenberger1976a;keener1993a;massey1983a;tobe1985a
|publication title=Syst. Nat. ed.
|publication year=1759
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/f6b125a955440c0872999024f038d74684f65921/coarse_grained_fna_xml/V3/V3_496.xml
|genus=Hydrastis
}}<!--

-->[[Category:Treatment]][[Category:Ranunculaceae]]