View source for Timmiella

{{Treatment/ID
|accepted_name=Timmiella
|accepted_authority=(De Notaris) Limpricht
|publications={{Treatment/Publication
|title=Laubm. Deutchl.
|place=1: 590. 1888 ,
}}
|basionyms={{Treatment/ID/Basionym
|name=Trichostomum sect. Timmiella
|authority=De Notaris
|publication_title=Comment. Soc. Crittog. Ital.
|publication_place=2: 100. 1866
}}
|synonyms=
|hierarchy=Pottiaceae;Pottiaceae subfam. Timmielloideae;Timmiella
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Pottiaceae]]</div><div class="higher-taxon"><small>subfamily</small>[[Pottiaceae subfam. Timmielloideae]]</div><div class="higher-taxon"><small>genus</small>[[Timmiella]]</div></div>
|etymology=Genus Timmia and Latin -ella, diminutive, alluding to resemblance
|volume=Volume 27
|mention_page=page 38, 477, 482
|treatment_page=page 481
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>loosely cespitose to cushion-forming, often rosulate, green, often dark distally, brown basally. <b>Stems</b> to 1.5 cm; hyalodermis present, weak, sclerodermis present in 1–3 layers, central strand very strong; axillary hairs of ca. 7 cells, all hyaline or proximal cell firm-walled. <b>Leaves</b> incurved, tubulose when dry, spreading when moist; long-elliptic to ligulate or broadly lanceolate, often wasp-waisted, adaxial surface broadly channeled across leaf, 3.5–5 mm; base often broadened, elliptic, somewhat sheathing, proximal margins occasionally with distinct shoulders; distal margins plane to weakly incurved, distinctly denticulate or serrulate to near base, usually strongly serrate near apex, lamina 2-stratose except 1-stratose along margins; apex acute; costa percurrent, tapering to apex and much broadened before mid leaf, adaxial outgrowths absent, adaxial cells quadrate and bulging in 6–19 rows; transverse section flattened, reniform or elliptic, adaxial epidermis present, 2-stratose or occasionally 3-stratose, adaxial stereid band present, guide cells 6–16 in 1 layer, hydroid strand present, often multiple, abaxial stereid band present, crescent-shaped in section, abaxial epidermis present; proximal cells differentiated straight across leaf, bulging rectangular, 10–18 µm wide, mostly 3–4:1, walls of proximal cells thin, hyaline to yellowish; distal medial cells quadrate to rounded-hexagonal, 9–12 µm wide, 1:1; papillae absent, cell walls mostly evenly thickened, lumina often rounded, bulging adaxially, cells 2-stratose but staggered (not situated directly over each other). <b>Specialized</b> asexual reproduction not seen. <b>Sexuality</b> dioicous or monoicous (autoicous, synoicous or apparently rhizautoicous). <b>Perichaetia</b> terminal, base of interior leaves often shortly and broadly clasping, scarcely different from cauline leaves. <b>Seta</b> ca. 0.8–2.5 cm. <b>Capsule</b> stegocarpous, theca long-elliptic to cylindric, occasionally somewhat ventricose, 2–4 mm, annulus absent or of 1–several layers of highly vesiculose cells, revoluble or deciduous in pieces; operculum long-conic to rostrate; peristome teeth of 32 filaments [16 paired teeth], filamentous [linear-lanceolate] in shape, twisted weakly to once twisted clockwise or straight, teeth to 300–700 µm [rudimentary]. <b>Calyptra</b> cucullate. <b>Spores</b> 8–13 µm. KOH laminal color reaction yellow.</span><!--

-->{{Treatment/Body
|distribution=North America;Mexico;Central America;South America;Europe;Asia;Africa;Pacific Islands.
|discussion=<p>Species 13 (2 in the flora).</p><!--
--><p>The salient characters of <i>Timmiella</i> are the laminal margins plane to weakly incurved, denticulate to dentate, the costa very wide, and the laminal cells are bulging adaxially but flat abaxially, and are medially 2-stratose but not aligned directly over each other near the costa. The opercular cells are in straight rows or twisted clockwise. Without sporophytes, collections are commonly difficult to name to species, and a revision worldwide is much needed. R. H. Zander (1993) placed it as phylogenetically basal to the <i>Pottiaceae</i>. Recent molecular work (O. Werner et al. 2004) indicated that the genus is phylogenetically distant from the <i>Pottiaceae</i>, but Zander (2006) justified retention in the <i>Pottiaceae</i> by appeal to Dollo’s Rule.</p>
|tables=
|references={{Treatment/Reference
|id=zander1994a
|text=Zander, R. H. 1994b. Timmiella. In: A. J. Sharp et al., eds. The moss flora of Mexico. Mem. New York Bot. Gard. 69: 243–246.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Leaf base narrower to little wider than the limb; autoicous or synoicous, seldom apparently dioicous; seta usually 1.5-2.5 cm, theca reaching 3-4 mm, operculum commonly 1-1.5 mm, peristome distinctly twisted
|[[Timmiella anomala|Timmiella anomala]]
|-id=key-0-1
|1
|Leaf base distinctly and rather abruptly wider than the limb; dioicous; seta 0.8-1.2 cm, theca usually 2-2.8 mm, operculum 0.6-1 mm, peristome not or slightly twisted
|[[Timmiella crassinervis|Timmiella crassinervis]]
|}
</div></div><!--

-->{{#Taxon:
name=Timmiella
|author=Richard H. Zander
|authority=(De Notaris) Limpricht
|rank=genus
|parent rank=subfamily
|synonyms=
|basionyms=Trichostomum sect. Timmiella
|family=Pottiaceae
|distribution=North America;Mexico;Central America;South America;Europe;Asia;Africa;Pacific Islands.
|reference=zander1994a
|publication title=Laubm. Deutchl.
|publication year=
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/f6b125a955440c0872999024f038d74684f65921/coarse_grained_fna_xml/V27/V27_696.xml
|subfamily=Pottiaceae subfam. Timmielloideae
|genus=Timmiella
}}<!--

-->[[Category:Treatment]][[Category:Pottiaceae subfam. Timmielloideae]]