View source for Cystopteris utahensis

{{Treatment/ID
|accepted_name=Cystopteris utahensis
|accepted_authority=Windham & Haufler in Haufler & Windham
|publications={{Treatment/Publication
|title=in Haufler & Windham,Amer. Fern J.
|place=81: 13. 1991
|year=1991
}}
|basionyms=
|synonyms=
|hierarchy=Dryopteridaceae;Cystopteris;Cystopteris utahensis
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Dryopteridaceae]]</div><div class="higher-taxon"><small>genus</small>[[Cystopteris]]</div><div class="higher-taxon"><small>species</small>[[Cystopteris utahensis]]</div></div>
|volume=Volume 2
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>creeping, not cordlike, internodes short, heavily beset with old petiole bases, hairs absent; scales lanceolate, ± clathrate, radial walls dark brown, thick, luminae clear. <b>Leaves</b> monomorphic, clustered at stem apex, to 45 cm, nearly all bearing sori. <b>Petiole</b> green to straw-colored throughout or darker near base, shorter than blade, base sparsely scaly. <b>Blade</b> deltate to narrowly deltate, 2-pinnate-pinnatifid, usually widest at or near base, apex short-attenuate; rachis and costae with unicellular, gland-tipped hairs, misshapen bulblets present or absent; axils of pinnae usually with multicellular, gland-tipped hairs. <b>Pinnae</b> typically perpendicular to rachis, not curving toward blade apex, margins serrate; proximal pinnae pinnatifid to pinnate-pinnatifid, ± equilateral, basiscopic pinnules not enlarged; basal basiscopic pinnules sessile or short-stalked, base truncate to obtuse, distal pinnae ovate to oblong. <b>Veins</b> directed into teeth and notches. <b>Indusia</b> cup-shaped, apex truncate, with scattered, unicellular, gland-tipped hairs. <b>Spores</b> spiny, usually 39–48 µm. <b>2n</b> = 168.</span><!--

-->{{Treatment/Body
|phenology=Sporulating summer–fall.
|habitat=Cracks and ledges on cliffs, on calcareous substrates including sandstone, limestone, and dacite
|elevation=1300–2700 m
|distribution=Ariz.;Colo.;Tex.;Utah.
|discussion=<p><i>Cystopteris utahensis</i> is an allopolyploid derived from the diploid species <i>C. bulbifera</i> and <i>C. reevesiana</i> (C. H. Haufler and M. D. Windham 1991). Because <i>C. utahensis</i> shares one parent (<i>C. bulbifera</i>) with <i>C. tennesseensis</i> and because of morphologic similarities between <i>C. reevesiana</i> and <i>C. protrusa</i> (the second diploid parent of <i>C. tennesseensis</i>), populations of <i>C. utahensis</i> were previously considered to have originated by long-distance dispersal from eastern populations of <i>C. tennesseensis</i>. Genetic studies using isozyme markers, however, indicated that <i>C. utahensis</i> was a distinct species and stimulated the discovery of morphologic criteria for distinguishing it from its eastern cousin. When combined with the geographic separation of the two tetraploids, the minor differences in indument features provide a means of circumscribing this genetically distinct species. Potential confusion in identifying <i>C. utahensis</i> arises because sterile triploid hybrids may form when it is sympatric with the more common diploid <i>C. reevesiana</i>.</p>
|tables=
|references=
}}<!--

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-->{{#Taxon:
name=Cystopteris utahensis
|author=
|authority=Windham & Haufler in Haufler & Windham
|rank=species
|parent rank=genus
|synonyms=
|basionyms=
|family=Dryopteridaceae
|phenology=Sporulating summer–fall.
|habitat=Cracks and ledges on cliffs, on calcareous substrates including sandstone, limestone, and dacite
|elevation=1300–2700 m
|distribution=Ariz.;Colo.;Tex.;Utah.
|reference=None
|publication title=in Haufler & Windham,Amer. Fern J.
|publication year=1991
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/f6b125a955440c0872999024f038d74684f65921/coarse_grained_fna_xml/V2/V2_501.xml
|genus=Cystopteris
|species=Cystopteris utahensis
}}<!--

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