View source for Woodsia cochisensis

{{Treatment/ID
|accepted_name=Woodsia cochisensis
|accepted_authority=Windham
|publications={{Treatment/Publication
|title=Contr. Univ. Michigan Herb.
|place=19: 54. 1993
|year=1993
}}
|common_names=Cochise cliff fern
|basionyms=
|synonyms=
|hierarchy=Dryopteridaceae;Woodsia;Woodsia cochisensis
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Dryopteridaceae]]</div><div class="higher-taxon"><small>genus</small>[[Woodsia]]</div><div class="higher-taxon"><small>species</small>[[Woodsia cochisensis]]</div></div>
|volume=Volume 2
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, erect to ascending, with a few persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. <b>Leaves</b> 5–25 × 1.5–6 cm. <b>Petiole</b> light brown or straw-colored throughout when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. <b>Blade</b> narrowly lanceolate to ovate, pinnate-pinnatifid to 2-pinnate proximally, sparsely to moderately glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with glandular hairs and occasional hairlike scales. <b>Pinnae</b> ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate; largest pinnae with 4–9 pairs of pinnules; abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales. <b>Pinnules</b> dentate, often shallowly lobed; margins lustrous adaxially, usually thickened, lacking cilia but sparsely glandular, with occasional 1–2-celled translucent projections. <b>Vein</b> tips enlarged to form whitish hydathodes visible adaxially. <b>Indusia</b> of relatively broad segments; segments multiseriate most of length, usually divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia. <b>Spores</b> averaging 43–49 µm. <b>2n</b> = 152.</span><!--

-->{{Treatment/Body
|phenology=Sporulating late spring–fall.
|habitat=Shaded ledges and alcoves near springs and seeps, usually on granitic or volcanic substrates
|elevation=1000–2200 m
|distribution=Ariz.;N.Mex.;n Mexico.
|discussion=<p><i>Woodsia cochisensis</i> traditionally has been identified as <i>W. plummerae</i> or (rarely) W. mexicana Fée. It is readily separated from <i>W. plummerae</i> by the characteristics given in the key, and from North American members of the mexicana group (<i>W. phillipsii</i> and <i>W. neomexicana</i>) by having indusial segments that are broad and nonfilamentous at the base. <i>Woodsia cochisensis</i> is less glandular than typical W. mexicana from northeastern Mexico and is further distinguished from that species by the thickened, lustrous pinnule margins and well-developed hydathodes. Isozyme and chromosome studies suggest that <i>W. cochisensis</i> is an allotetraploid that may have originated through hybridization between <i>W. phillipsii</i> and an undescribed Mexican diploid (M. D. Windham 1993). It crosses with the former species to produce sterile triploids of intermediate morphology.</p>
|tables=
|references=
}}<!--

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-->{{#Taxon:
name=Woodsia cochisensis
|author=
|authority=Windham
|rank=species
|parent rank=genus
|synonyms=
|basionyms=
|family=Dryopteridaceae
|phenology=Sporulating late spring–fall.
|habitat=Shaded ledges and alcoves near springs and seeps, usually on granitic or volcanic substrates
|elevation=1000–2200 m
|distribution=Ariz.;N.Mex.;n Mexico.
|reference=None
|publication title=Contr. Univ. Michigan Herb.
|publication year=1993
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/f6b125a955440c0872999024f038d74684f65921/coarse_grained_fna_xml/V2/V2_370.xml
|genus=Woodsia
|species=Woodsia cochisensis
}}<!--

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