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Roxanne I. Hastings +  and Henk C. Greven +
unknown +
North America +, Mexico +, Central America +, South America +, Eurasia +, Africa +, Pacific Islands +, Australia +  and Antarctic. +
Sp. Musc. Frond., +
Dryptodon +  and Hydrogrimmia +
Grimmia subg. Undetermined +
subgenus +