View source for Drymaria

{{Treatment/ID
|accepted_name=Drymaria
|accepted_authority=Willdenow ex Schultes in J. J. Roemer et al.
|publications={{Treatment/Publication
|title=in J. J. Roemer et al.,Syst. Veg.
|place=5: xxxi, 406. 1819
|year=1819
}}
|common_names=Drymary
|basionyms=
|synonyms=
|hierarchy=Caryophyllaceae;Caryophyllaceae subfam. Polycarpoideae;Drymaria
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Caryophyllaceae]]</div><div class="higher-taxon"><small>subfamily</small>[[Caryophyllaceae subfam. Polycarpoideae]]</div><div class="higher-taxon"><small>genus</small>[[Drymaria]]</div></div>
|etymology=Greek drymos, forest, alluding to habitat of at least one species
|volume=Volume 5
|mention_page=page 5, 7, 10
|treatment_page=page 9
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism duration;whole_organism duration;whole_organism growth form;caudex architecture"><b>Herbs,</b> annual or perennial, caudices often branched.</span> <span class="statement" id="st-d0_s1" data-properties="taproot size;taproot shape"><b>Taproots </b>slender, elongate.</span> <span class="statement" id="st-d0_s2" data-properties="stem orientation;stem architecture;stem architecture;stem shape"><b>Stems </b>sprawling to erect, simple or branching proximally or throughout, terete.</span> <span class="statement" id="st-d0_s3" data-properties="leaf arrangement;leaf arrangement;leaf arrangement;leaf fusion;leaf architecture;leaf architecture;leaf architecture;line texture;line size or width"><b>Leaves </b>opposite or appearing whorled, connate by membranous to thickened line, petiolate or sessile, stipulate (D. pachyphylla not stipulate);</span> <span class="statement" id="st-d0_s4" data-properties="stipule quantity;stipule coloration;stipule architecture;stipule shape;stipule shape;stipule size;margin architecture or shape;apex shape">stipules 2 per node, white to tan, simple or divided into segments, subulate to filiform, often minute, margins entire, apex acute to acuminate;</span> <span class="statement" id="st-d0_s5" data-properties="blade architecture;blade shape;blade shape;blade shape;blade shape;blade shape;blade texture;apex shape">blade 1–5-veined, linear to lanceolate, spatulate, ovate, reniform, or orbiculate, not succulent, apex rounded to acuminate.</span> <span class="statement" id="st-d0_s6" data-properties="inflorescence position;inflorescence position;inflorescence architecture;cyme architecture;cyme architecture;cyme arrangement;flower architecture or arrangement or growth form;flower position"><b>Inflorescences </b>terminal or axillary, open to congested, bracteate cymes or umbelliform clusters or flowers solitary, axillary;</span> <span class="statement" id="st-d0_s7" data-properties="bract arrangement;bract texture;central portion growth form or texture">bracts paired, scarious or central portion herbaceous.</span> <span class="statement" id="st-d0_s8" data-properties="pedicel orientation"><b>Pedicels </b>erect to spreading or reflexed.</span> <span class="statement" id="st-d0_s9" data-properties="perianth position;androecium position"><b>Flowers:</b> perianth and androecium hypogynous;</span> <span class="statement" id="st-d0_s10" data-properties="sepal quantity;sepal fusion;sepal coloration;sepal shape;sepal shape;sepal shape;sepal atypical some measurement;sepal some measurement;sepal growth form or texture;margin coloration;margin texture;apex shape;apex architecture;apex architecture">sepals 5, distinct, white, lanceolate to oblong, ovate, or orbiculate, 1.5–4.8 (–5) mm, herbaceous, margins white to purple, scarious, apex acuminate to rounded, hooded or not;</span> <span class="statement" id="st-d0_s11" data-properties="petal atypical quantity;petal quantity;petal presence;petal coloration;claw size or width;claw shape;claw shape;trunk shape;trunk size;trunk architecture;trunk shape;auricle presence;blade apex shape">petals (3–) 5, sometimes absent, white, claw narrow, tapering distally or with oblong or expanded, sessile or short-clawed trunk, auricles absent, blade apex divided into 2 or 4 lobes;</span> <span class="statement" id="st-d0_s12" data-properties="filament arrangement;sepal arrangement">nectaries at base of filaments opposite sepals;</span> <span class="statement" id="st-d0_s13" data-properties="stamen quantity">stamens 5;</span> <span class="statement" id="st-d0_s14" data-properties="filament fusion;filament fusion">filaments distinct or briefly connate proximally;</span> <span class="statement" id="st-d0_s15" data-properties="style quantity;style quantity;style fusion;style length;style fusion;style shape;style some measurement;style pubescence">styles 3, occasionally 2, connate proximally for 1/2 of length, rarely to nearly distinct (D. cordata), filiform, 0.1–0.3 mm, glabrous proximally;</span> <span class="statement" id="st-d0_s16" data-properties="stigma quantity;stigma quantity;stigma arrangement or course or shape;stigma relief;stigma quantity">stigmas 3, occasionally 2, linear along adaxial surfaces of styles (or branches), obscurely papillate (30×).</span> <span class="statement" id="st-d0_s17" data-properties="capsule shape;valve atypical quantity;valve quantity;valve orientation"><b>Capsules </b>ellipsoid to globose, opening by (2–) 3 spreading to recurved valves;</span> <span class="statement" id="st-d0_s18" data-properties="carpophore presence">carpophore absent.</span> <span class="statement" id="st-d0_s19" data-properties="seed quantity;seed coloration;seed coloration;seed coloration;seed coloration;seed coloration;seed coloration;seed coloration;seed shape;seed shape;seed shape;seed shape;seed relief;marginal wing presence"><b>Seeds </b>3–25, tan, reddish-brown, dark-brown, black, or transparent (white embryo visible), horseshoe, snail-shell or teardrop-shaped, compressed laterally, at least somewhat, tuberculate, marginal wing absent, appendage absent.</span> <span class="statement" id="st-d0_s20" data-properties="appendage presence;x chromosome atypical quantity;x chromosome quantity">x = (11), 12.</span><!--

-->{{Treatment/Body
|distribution=sw United States;Mexico;West Indies;Central America;South America;in Asia (Indonesia);e;s Africa;Australia;Pacific Islands
|discussion=<p>Species 48 (9 in the flora).</p><!--
--><p>Drymaria arenarioides Willdenow ex Schultes, alfombrilla, is on the Federal Noxious Weed List. It is highly toxic to livestock and is native in northwestern Mexico, where it has been reported within a few miles of the United States border.</p><!--
--><p>J. A. Duke (1961) proposed an infraspecific classification for Drymaria consisting of 17 “informal” series that he did not validly publish. Some of Duke’s series were cited by M. Escamilla and V. Sosa (2000), apparently assuming that they were “real” series.</p>
|tables=
|references={{Treatment/Reference
|id=duke1961a
|text=Duke, J. A. 1961. Preliminary revision of the genus Drymaria. Ann. Missouri Bot. Gard. 48: 173–268.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Blades of cauline leaves ovate, deltate, cordate, reniform, orbiculate, or suborbiculate, 4-25 mm wide, base obtuse or cordate to rounded
|[[#key-0-2| > 2]]
|-id=key-0-1
|1
|Blades of cauline leaves linear to oblong or lanceolate, 0.2-3 mm wide, base briefly obtuse to attenuate
|[[#key-0-5| > 5]]
|-id=key-0-2
|2
|Stems erect or ascending; petals 2-fid, lobe apex not notched; seeds 0.5-0.7 mm
|[[Drymaria glandulosa|Drymaria glandulosa]]
|-id=key-0-2
|2
|Stems prostrate or sprawling; petals 2- or 4-fid, if 2-fid, lobe apex distinctly notched or seeds 1-1.5 mm
|[[#key-0-3| > 3]]
|-id=key-0-3
|3
|Plants glabrous, glaucous, succulent; petals 4-fid; leaves appearing whorled, not stipulate; inflorescences umbelliform clusters
|[[Drymaria pachyphylla|Drymaria pachyphylla]]
|-id=key-0-3
|3
|Plants often glandular in inflorescence, not glaucous, herbaceous; petals 2-fid; leaves opposite, stipules present or deciduous in part; inflorescences open cymes
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Petals with lobe apex rounded to acute, 1-veined, vein not branched; seeds 1-1.5 mm, tubercles rounded
|[[Drymaria cordata|Drymaria cordata]]
|-id=key-0-4
|4
|Petals with lobe apex distinctly notched, 1-veined, vein dichotomously branched;seeds 0.5-0.7 mm, tubercles conic
|[[Drymaria laxiflora|Drymaria laxiflora]]
|-id=key-0-5
|5
|Cauline leaves appearing whorled, at least in part
|[[#key-0-6| > 6]]
|-id=key-0-5
|5
|Cauline leaves opposite
|[[#key-0-7| > 7]]
|-id=key-0-6
|6
|Plants viscid; stems prostrate; sepals stipitate-glandular; inflorescences 4-7-flowered, axillary and terminal cymes
|[[Drymaria viscosa|Drymaria viscosa]]
|-id=key-0-6
|6
|Plants glabrous or sparsely glandular; stems erect; sepals glabrous; inflorescences 3-30+-flowered, terminal cymes initially becoming racemose
|[[Drymaria molluginea|Drymaria molluginea]]
|-id=key-0-7
|7
|Petals 11/ 1/ 2 times as long as sepals
|[[Drymaria effusa|Drymaria effusa]]
|-id=key-0-7
|7
|Petals 1/ 2-1 times as long as sepals
|[[#key-0-8| > 8]]
|-id=key-0-8
|8
|Herbaceous portion of sepals ± oblong, apex blunt or rounded, veins ± parallel, apically confluent
|[[Drymaria depressa|Drymaria depressa]]
|-id=key-0-8
|8
|Herbaceous portion of sepals lanceolate, apex acute to acuminate, veins with lateral pair distinctly arcing outward
|[[Drymaria leptophylla|Drymaria leptophylla]]
|}
</div></div><!--

-->{{#Taxon:
name=Drymaria
|author=Ronald L. Hartman
|authority=Willdenow ex Schultes in J. J. Roemer et al.
|rank=genus
|parent rank=subfamily
|synonyms=
|basionyms=
|family=Caryophyllaceae
|illustrator=Barbara Alongi
|illustration copyright=Flora of North America Association
|distribution=sw United States;Mexico;West Indies;Central America;South America;in Asia (Indonesia);e;s Africa;Australia;Pacific Islands
|reference=duke1961a
|publication title=in J. J. Roemer et al.,Syst. Veg.
|publication year=1819
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V5/V5_2.xml
|subfamily=Caryophyllaceae subfam. Polycarpoideae
|genus=Drymaria
|androecium position=hypogynous
|apex architecture=not;hooded
|apex shape=acuminate;rounded
|appendage presence=absent
|auricle presence=absent
|blade apex shape=divided
|blade architecture=1-5-veined
|blade shape=linear;lanceolate spatulate ovate reniform or orbiculate
|blade texture=not succulent
|bract arrangement=paired
|bract texture=scarious
|capsule shape=ellipsoid;globose
|carpophore presence=absent
|caudex architecture=branched
|central portion growth form or texture=herbaceous
|claw shape=with oblong or expanded , sessile or short-clawed trunk;tapering
|claw size or width=narrow
|cyme architecture=umbelliform;bracteate
|cyme arrangement=cluster
|filament arrangement=opposite
|filament fusion=connate;distinct
|flower architecture or arrangement or growth form=solitary
|flower position=axillary
|inflorescence architecture=open;congested
|inflorescence position=axillary;terminal
|leaf architecture=stipulate;sessile;petiolate
|leaf arrangement=whorled;appearing;opposite
|leaf fusion=connate
|line size or width=thickened
|line texture=membranous
|margin architecture or shape=entire
|margin coloration=white;purple
|margin texture=scarious
|marginal wing presence=absent
|pedicel orientation=erect;spreading or reflexed
|perianth position=hypogynous
|petal atypical quantity=3;5
|petal coloration=white
|petal presence=absent
|petal quantity=5
|seed coloration=transparent;black;transparent;black;dark-brown;reddish-brown;tan
|seed quantity=3;25
|seed relief=tuberculate
|seed shape=compressed;teardrop--shaped;snail-shell;horseshoe
|sepal arrangement=opposite
|sepal atypical some measurement=4.8mm;5mm
|sepal coloration=white
|sepal fusion=distinct
|sepal growth form or texture=herbaceous
|sepal quantity=5
|sepal shape=lanceolate;oblong ovate or orbiculate
|sepal some measurement=1.5mm;4.8mm
|stamen quantity=5
|stem architecture=branching;simple
|stem orientation=sprawling;erect
|stem shape=terete
|stigma arrangement or course or shape=linear
|stigma quantity=[30;2;3
|stigma relief=papillate
|stipule architecture=simple
|stipule coloration=white;tan
|stipule quantity=2
|stipule shape=subulate;filiform
|stipule size=minute
|style fusion=distinct;connate
|style length=1/2
|style pubescence=glabrous
|style quantity=2;3
|style shape=filiform
|style some measurement=0.1mm;0.3mm
|taproot shape=elongate
|taproot size=slender
|trunk architecture=sessile
|trunk shape=short-clawed;oblong
|trunk size=expanded
|valve atypical quantity=2;3
|valve orientation=spreading;recurved
|valve quantity=3
|whole_organism duration=perennial;annual
|whole_organism growth form=herb
|x chromosome atypical quantity=11
|x chromosome quantity=12
}}<!--

-->[[Category:Treatment]][[Category:Caryophyllaceae subfam. Polycarpoideae]]