Difference between revisions of "Ostrya"

Scopoli

Fl. Carniol., 414. 1760, name conserved.

Common names: Hop-hornbeam
Etymology: Latin ostrya, hop-hornbeam, from Greek ostryos, scale, in reference to the scaly infructescences
Treatment appears in FNA Volume 3.
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--><span class="statement" id="st-d0_s0" data-properties="tree some measurement"><b>Trees,</b> 9–18 m;</span> <span class="statement" id="st-d0_s1" data-properties="trunk quantity;trunk architecture;trunk architecture or texture;trunk shape;branch shape">trunks usually 1, branching mostly deliquescent, trunk and branches terete.</span> <span class="statement" id="st-d0_s2" data-properties="bark coloration;bark coloration;bark coloration;bark width;bark architecture or pubescence or relief;strip pubescence;strip orientation;scale pubescence;scale orientation"><b>Bark </b>of trunk and branches brownish gray to light-brown, thin, smooth, breaking and shredding into shaggy vertical strips and scales;</span> <span class="statement" id="st-d0_s3" data-properties="lenticel prominence">lenticels generally inconspicuous.</span> <span class="statement" id="st-d0_s4" data-properties="wood coloration;wood coloration;wood coloration;wood texture;wood texture"><b>Wood </b>nearly white to light-brown, very hard and heavy, texture fine.</span> <span class="statement" id="st-d0_s5" data-properties="twig arrangement"><b>Branches,</b> branchlets, and twigs conspicuously 2-ranked;</span> <span class="statement" id="st-d0_s6" data-properties="twig life cycle;twig variability;short-shoot length or size">young twigs differentiated into long and short-shoots.</span> <span class="statement" id="st-d0_s7" data-properties="bud season;bud architecture;bud shape;bud shape;apex shape"><b>Winter </b>buds sessile, ovoid, somewhat laterally compressed, apex acute;</span> <span class="statement" id="st-d0_s8" data-properties="scale quantity;scale arrangement;scale coloration or pubescence or relief">scales many, imbricate, longitudinally striate.</span> <span class="statement" id="st-d0_s9" data-properties="leaf arrangement;short-shoot length or size"><b>Leaves </b>on long and short-shoots, 2-ranked.</span> <span class="statement" id="st-d0_s10" data-properties="leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade shape;leaf-blade length;leaf-blade width;lateral-vein width;margin architecture or shape;margin architecture or shape;margin architecture or shape"><b>Leaf-</b>blade narrowly ovate to ovate, elliptic, or obovate with 10 or more pairs of lateral-veins, 2.5–13 × 1.5–6 cm, thin, margins doubly serrate to serrulate;</span> <span class="statement" id="st-d0_s11" data-properties="surface pubescence;surface pubescence;surface pubescence">surfaces abaxially glabrous to tomentose.</span> <span class="statement" id="st-d0_s12" data-properties="catkin architecture;catkin position or structure subtype;catkin prominence;catkin size;season growth order"><b>Inflorescences:</b> staminate catkins terminal on branches, mostly in small, racemose clusters, formed previous growing season and exposed during winter, expanding with leaves;</span> <span class="statement" id="st-d0_s13" data-properties="catkin architecture;catkin position;catkin architecture;catkin architecture or arrangement or growth form;catkin orientation;catkin shape;growth height or length or size;growth position;growth architecture;growth life cycle;beak arrangement or density;flower arrangement or density">pistillate catkins proximal to staminate on short, lateral, leafy new growth, solitary, ± erect, elongate, bracts and flowers uncrowded.</span> <span class="statement" id="st-d0_s14" data-properties="flower architecture;flower arrangement;catkin quantity;receptacle pubescence"><b>Staminate </b>flowers in catkins 3 per bract, crowded together on pilose receptacle;</span> <span class="statement" id="st-d0_s15" data-properties="stamen atypical quantity;stamen quantity;stamen height or length or size">stamens 3 (–6), short;</span> <span class="statement" id="st-d0_s16" data-properties="filament shape;filament dehiscence">filaments often divided part-way to base;</span> <span class="statement" id="st-d0_s17" data-properties="anther shape;anther architecture or structure in adjective form;part quantity;apex pubescence">anthers divided into 2 parts, each 1-locular, apex pilose.</span> <span class="statement" id="st-d0_s18" data-properties="flower architecture;flower quantity"><b>Pistillate </b>flowers 2 per bract.</span> <span class="statement" id="st-d0_s19" data-properties="beak condition;beak shape"><b>Infructescences </b>loosely imbricate, strobiloid clusters of closed inflated bracts;</span> <span class="statement" id="st-d0_s20" data-properties="infructescence arrangement;infructescence architecture;infructescence arrangement;infructescence arrangement;infructescence orientation;infructescence shape">clusters pendulous, elongate;</span> <span class="statement" id="st-d0_s21" data-properties="beak duration;beak shape;beak shape;fruit quantity">bracts deciduous with fruit, inflated, bladderlike, each bract enclosing 1 fruit.</span> <span class="statement" id="st-d0_s22" data-properties="nutlet size;sepal duration"><b>Fruits </b>small nutlets, ovoid, longitudinally ribbed, often crowned with persistent sepals and styles.</span> <span class="statement" id="st-d0_s23" data-properties="fruit shape;fruit architecture or shape;fruit architecture;x chromosome quantity">x = 8.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees,</b> 9–18 m; trunks usually 1, branching mostly deliquescent, trunk and branches terete. <b>Bark</b> of trunk and branches brownish gray to light brown, thin, smooth, breaking and shredding into shaggy vertical strips and scales; lenticels generally inconspicuous. <b>Wood</b> nearly white to light brown, very hard and heavy, texture fine. <b>Branches</b>, branchlets, and twigs conspicuously 2-ranked; young twigs differentiated into long and short shoots. <b>Winter</b> buds sessile, ovoid, somewhat laterally compressed, apex acute; scales many, imbricate, longitudinally striate. <b>Leaves</b> on long and short shoots, 2-ranked. <b>Leaf</b> blade narrowly ovate to ovate, elliptic, or obovate with 10 or more pairs of lateral veins, 2.5–13 × 1.5–6 cm, thin, margins doubly serrate to serrulate; surfaces abaxially glabrous to tomentose. <b>Inflorescences</b>: staminate catkins terminal on branches, mostly in small, racemose clusters, formed previous growing season and exposed during winter, expanding with leaves; pistillate catkins proximal to staminate on short, lateral, leafy new growth, solitary, ± erect, elongate, bracts and flowers uncrowded. <b>Staminate</b> flowers in catkins 3 per bract, crowded together on pilose receptacle; stamens 3(–6), short; filaments often divided part way to base; anthers divided into 2 parts, each 1-locular, apex pilose. <b>Pistillate</b> flowers 2 per bract. <b>Infructescences</b> loosely imbricate, strobiloid clusters of closed inflated bracts; clusters pendulous, elongate; bracts deciduous with fruit, inflated, bladderlike, each bract enclosing 1 fruit. <b>Fruits</b> small nutlets, ovoid, longitudinally ribbed, often crowned with persistent sepals and styles. <b>x</b> = 8.</span><!--
  
 
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|publication year=
 
|publication year=
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_3.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V3/V3_3.xml
 
|subfamily=Betulaceae subfam. Coryloideae
 
|subfamily=Betulaceae subfam. Coryloideae
 
|genus=Ostrya
 
|genus=Ostrya
|anther architecture or structure in adjective form=1-locular
 
|anther shape=divided
 
|apex pubescence=pilose
 
|apex shape=acute
 
|bark architecture or pubescence or relief=smooth
 
|bark coloration=brownish gray;light-brown
 
|bark width=thin
 
|beak arrangement or density=uncrowded
 
|beak condition=closed
 
|beak duration=deciduous
 
|beak shape=bladderlike;inflated;inflated
 
|branch shape=terete
 
|bud architecture=sessile
 
|bud season=winter
 
|bud shape=compressed;ovoid
 
|catkin architecture=staminate;pistillate;staminate
 
|catkin architecture or arrangement or growth form=solitary
 
|catkin orientation=erect
 
|catkin position=proximal
 
|catkin position or structure subtype=terminal
 
|catkin prominence=exposed
 
|catkin quantity=3
 
|catkin shape=elongate
 
|catkin size=expanding
 
|filament dehiscence=part-way
 
|filament shape=divided
 
|flower architecture=pistillate;staminate
 
|flower arrangement=crowded
 
|flower arrangement or density=uncrowded
 
|flower quantity=2
 
|fruit architecture=crowned
 
|fruit architecture or shape=ribbed
 
|fruit quantity=1
 
|fruit shape=ovoid
 
|growth architecture=leafy
 
|growth height or length or size=short
 
|growth life cycle=new
 
|growth position=lateral
 
|infructescence architecture=strobiloid
 
|infructescence arrangement=cluster;cluster;imbricate
 
|infructescence orientation=pendulous
 
|infructescence shape=elongate
 
|lateral-vein width=1.5cm;6cm
 
|leaf arrangement=2-ranked
 
|leaf-blade length=2.5cm;13cm
 
|leaf-blade shape=narrowly ovate;ovate elliptic or obovate
 
|leaf-blade width=thin
 
|lenticel prominence=inconspicuous
 
|margin architecture or shape=doubly serrate;serrulate
 
|nutlet size=small
 
|part quantity=2
 
|receptacle pubescence=pilose
 
|scale arrangement=imbricate
 
|scale coloration or pubescence or relief=striate
 
|scale orientation=vertical
 
|scale pubescence=shaggy
 
|scale quantity=many
 
|season growth order=previous
 
|sepal duration=persistent
 
|short-shoot length or size=long;long
 
|stamen atypical quantity=3;6
 
|stamen height or length or size=short
 
|stamen quantity=3
 
|strip orientation=vertical
 
|strip pubescence=shaggy
 
|surface pubescence=abaxially glabrous;tomentose
 
|tree some measurement=9m;18m
 
|trunk architecture=branching
 
|trunk architecture or texture=deliquescent
 
|trunk quantity=1
 
|trunk shape=terete
 
|twig arrangement=2-ranked
 
|twig life cycle=young
 
|twig variability=differentiated
 
|wood coloration=nearly white;light-brown
 
|wood texture=fine;hard
 
|x chromosome quantity=8
 
 
}}<!--
 
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Revision as of 13:37, 27 July 2019

Trees, 9–18 m; trunks usually 1, branching mostly deliquescent, trunk and branches terete. Bark of trunk and branches brownish gray to light brown, thin, smooth, breaking and shredding into shaggy vertical strips and scales; lenticels generally inconspicuous. Wood nearly white to light brown, very hard and heavy, texture fine. Branches, branchlets, and twigs conspicuously 2-ranked; young twigs differentiated into long and short shoots. Winter buds sessile, ovoid, somewhat laterally compressed, apex acute; scales many, imbricate, longitudinally striate. Leaves on long and short shoots, 2-ranked. Leaf blade narrowly ovate to ovate, elliptic, or obovate with 10 or more pairs of lateral veins, 2.5–13 × 1.5–6 cm, thin, margins doubly serrate to serrulate; surfaces abaxially glabrous to tomentose. Inflorescences: staminate catkins terminal on branches, mostly in small, racemose clusters, formed previous growing season and exposed during winter, expanding with leaves; pistillate catkins proximal to staminate on short, lateral, leafy new growth, solitary, ± erect, elongate, bracts and flowers uncrowded. Staminate flowers in catkins 3 per bract, crowded together on pilose receptacle; stamens 3(–6), short; filaments often divided part way to base; anthers divided into 2 parts, each 1-locular, apex pilose. Pistillate flowers 2 per bract. Infructescences loosely imbricate, strobiloid clusters of closed inflated bracts; clusters pendulous, elongate; bracts deciduous with fruit, inflated, bladderlike, each bract enclosing 1 fruit. Fruits small nutlets, ovoid, longitudinally ribbed, often crowned with persistent sepals and styles. x = 8.

Distribution

Mostly north temperate zones

Discussion

Species ca. 5 (3 in the flora).

In North America Ostrya consists of small trees in the northern temperate deciduous forest zone and in the mountains of southwestern United States and adjacent Mexico. Mexican populations have generally been treated as conspecific with O. virginiana of eastern United States and Canada. They differ in various respects, however, including leaf shape and indumentum; the morphologic variation and phytogeography of the complex as a whole should be carefully examined. Ostrya carpinifolia Scopoli is a common and important forest tree of southern Europe.

Ostrya shares many features with Carpinus. The staminate catkins in most species of Ostrya are produced the season before anthesis but, unlike Carpinus, they are exposed during the winter. Dispersal occurs as it does in Carpinus, except that the bracts form closed, bladderlike structures rather than flat wings.

The wood of Ostrya is used for fuel, fence posts, and various other purposes. It was formerly utilized for manufacturing items subject to prolonged friction, including sleigh runners, wheel rims, and airplane propellers. Because of its hardness, it has been used for tool handles, mallet heads, and other hard wooden objects.

Key

1 Leaf blade (5–)8–10(–13) cm, apex usually abruptly acuminate; infructescences 3.5–6.5 cm; e, nc United States, adjacent Canada. Ostrya virginiana
1 Leaf blade 2.5–6.5 cm, apex acute, obtuse, or rounded; infructescences 2–4 cm; sw United States. > 2
2 Leaf blade ovate or broadly ovate-elliptic to broadly elliptic to nearly orbiculate; petiole and young twigs often bearing stipitate glands; staminate catkins 2–3 cm; w Texas, New Mexico, Arizona, Utah. Ostrya knowltonii
2 Leaf blade elliptic to elliptic-lanceolate; petiole and young twigs without stipitate glands; staminate catkins 3.5–5 cm; endemic to Chisos Mountains in Big Bend National Park, Texas. Ostrya chisosensis