familyPinaceae
genusAbies

Difference between revisions of "Abies bifolia"

A. Murray bis

Proc. Roy. Hort. Soc. London 3: 320. 1863.

Common names: Rocky Mountain alpine fir Rocky Mountain subalpine fir corkbark fir
Endemic
Synonyms: Abies subalpina Engelmann
Treatment appears in FNA Volume 2.
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|common_names=Rocky Mountain alpine fir;Rocky Mountain subalpine fir;corkbark fir
 
|common_names=Rocky Mountain alpine fir;Rocky Mountain subalpine fir;corkbark fir
 +
|special_status={{Treatment/ID/Special_status
 +
|code=E
 +
|label=Endemic
 +
}}
 
|basionyms=
 
|basionyms=
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Abies subalpina
 
|name=Abies subalpina
 
|authority=Engelmann
 
|authority=Engelmann
 +
|rank=species
 
}}
 
}}
 
|hierarchy=Pinaceae;Abies;Abies bifolia
 
|hierarchy=Pinaceae;Abies;Abies bifolia
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--><span class="statement" id="st-d0_s0" data-properties="tree some measurement"><b>Trees </b>to 30m;</span> <span class="statement" id="st-d0_s1" data-properties="trunk diameter">trunk to 0.45m diam.;</span> <span class="statement" id="st-d0_s2" data-properties="crown shape">crown spirelike.</span> <span class="statement" id="st-d0_s3" data-properties="bark coloration;bark width;bark architecture or pubescence or relief;bark architecture or pubescence;age architecture"><b>Bark </b>gray, thin, smooth, with age somewhat furrowed and scaly (toward southern end of range bark corky [corkbark fir]).</span> <span class="statement" id="st-d0_s4" data-properties="branch orientation;branch fragility or size;branch fragility"><b>Branches </b>diverging from trunk at right angles, stout, stiff;</span> <span class="statement" id="st-d0_s5" data-properties="twig arrangement;twig coloration;twig pubescence;twig coloration">twigs opposite to whorled, grayish, pubescence sparse, light-brown;</span> <span class="statement" id="st-d0_s6" data-properties="leaf-scar condition;periderm coloration">fresh leaf-scars with light-brown periderm.</span> <span class="statement" id="st-d0_s7" data-properties="bud prominence;bud coloration;bud shape;bud size;bud coating;apex shape"><b>Buds </b>exposed, brown, globose, small, resinous, apex rounded;</span> <span class="statement" id="st-d0_s8" data-properties="basal scale length or size;basal scale size or width;basal scale shape;basal scale shape;basal scale shape;basal scale pubescence;basal scale coating;basal scale coating;margin shape;margin shape;margin shape;apex shape;apex shape">basal scales long, narrow, isosceles triangular to spatulate, glabrous, resinous or not resinous, margins entire to rarely crenate, apex sharp-pointed or rounded.</span> <span class="statement" id="st-d0_s9" data-properties="leaf length;leaf width;leaf arrangement;leaf orientation;leaf fragility"><b>Leaves </b>1.1–2.5cm × 1.25–1.5mm, spiraled and turned upward, flexible;</span> <span class="statement" id="st-d0_s10" data-properties="">cross-section flat, grooved adaxially, sometimes only slightly so;</span> <span class="statement" id="st-d0_s11" data-properties="cross-section prominence or shape;cross-section architecture;cross-section odor">odor camphorlike;</span> <span class="statement" id="st-d0_s12" data-properties="stomatal count;row count">abaxial surface with 3–5 stomatal rows on each side of midrib;</span> <span class="statement" id="st-d0_s13" data-properties="adaxial surface coloration;adaxial surface coloration;adaxial surface coloration;adaxial surface pubescence;stomatal count;row count;row architecture;row count">adaxial surface light green to bluish green, usually glaucous, with 3–6 stomatal rows at midleaf, rows usually continuous to leaf base, usually more numerous toward leaf apex;</span> <span class="statement" id="st-d0_s14" data-properties="apex shape;apex shape;apex shape">apex slightly notched to rounded;</span> <span class="statement" id="st-d0_s15" data-properties="resin canal size;resin canal position;margin position">resin canals large, ± median, away from margins and midway between abaxial and adaxial epidermal layers.</span> <span class="statement" id="st-d0_s16" data-properties="pollination coloration"><b>Pollen </b>cones at pollination purplish.</span> <span class="statement" id="st-d0_s17" data-properties="seed-cone shape;seed-cone length;seed-cone width;seed-cone coloration;seed-cone coloration;seed-cone coloration;seed-cone architecture;apex shape"><b>Seed-</b>cones cylindric, 5–10 × 3–3.5cm, dark purple-blue to grayish purple, sessile, apex rounded;</span> <span class="statement" id="st-d0_s18" data-properties="scale length;scale width;scale pubescence">scales ca. 1.5 × 2.5cm, densely pubescent;</span> <span class="statement" id="st-d0_s19" data-properties="bract position">bracts included.</span> <span class="statement" id="st-d0_s20" data-properties="seed length;seed width;body coloration"><b>Seeds </b>5–7 × 2–3mm, body brown;</span> <span class="statement" id="st-d0_s21" data-properties="wing length;wing coloration">wing about 1.5 times as long as body, grayish brown;</span> <span class="statement" id="st-d0_s22" data-properties="cotyledon count">cotyledons 3–6.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees </b>to 30m; trunk to 0.45m diam.; crown spirelike. <b>Bark</b> gray, thin, smooth, with age somewhat furrowed and scaly (toward southern end of range bark is corky [corkbark fir]). <b>Branches</b> diverging from trunk at right angles, stout, stiff; twigs opposite to whorled, grayish, pubescence sparse, light brown; fresh leaf scars with light brown periderm. <b>Buds</b> exposed, brown, globose, small, resinous, apex rounded; basal scales long, narrow, isosceles triangular to spatulate, glabrous, resinous or not resinous, margins entire to rarely crenate, apex sharp-pointed or rounded. <b>Leaves</b> 1.1–2.5cm × 1.25–1.5mm, spiraled and turned upward, flexible; cross section flat, grooved adaxially, sometimes only slightly so; odor camphorlike; abaxial surface with 3–5 stomatal rows on each side of midrib; adaxial surface light green to bluish green, usually glaucous, with 3–6 stomatal rows at midleaf, rows usually continuous to leaf base, usually more numerous toward leaf apex; apex slightly notched to rounded; resin canals large, ± median, away from margins and midway between abaxial and adaxial epidermal layers. <b>Pollen</b> cones at pollination purplish. <b>Seed</b> cones cylindric, 5–10 × 3–3.5cm, dark purple-blue to grayish purple, sessile, apex rounded; scales ca. 1.5 × 2.5cm, densely pubescent; bracts included. <b>Seeds</b> 5–7 × 2–3mm, body brown; wing about 1.5 times as long as body, grayish brown; cotyledons 3–6.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
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|elevation=600–3600m
 
|elevation=600–3600m
 
|distribution=Alta.;B.C.;N.W.T.;Yukon;Ariz.;Colo.;Idaho;Mont.;Nev.;N.Mex.;Oreg.;Utah;Wash.;Wyo.
 
|distribution=Alta.;B.C.;N.W.T.;Yukon;Ariz.;Colo.;Idaho;Mont.;Nev.;N.Mex.;Oreg.;Utah;Wash.;Wyo.
|discussion=<p>Abies bifolia has been—and by many workers still is—included in synonymy under A. lasiocarpa or A. subalpina since about 1890, and A. subalpina under A. lasiocarpa since about the 1920s. Abies bifolia is distinct from A. lasiocarpa, however, in chemical tests on wood (H.S. Fraser and E.P. Swan 1972), lack of crystals in the ray parenchyma (R.W. Kennedy et al. 1968), lack of lasiocarpenonol (J.F. Manville and A.S. Tracey 1989), and distinct terpene patterns (R.S. Hunt and E.von Rudloff 1979). Abies bifolia also tends to have slightly shorter and fewer prominently notched leaves than A. lasiocarpa. The two are clearly separated by the color of their periderm and by the shape of their basal bud scales. These firs may be more distinct than the pairs A. balsamea -- A. fraseri and A. procera -- A. magnifica. A north-south transect, however, from south central Yukon to northern Washington yielded introgressed trees possessing characteristics of both A. lasiocarpa and A. bifolia, recalling the interior spruce (Canadian Forestry Service 1983), which has characteristics of both Picea glauca and P. engelmannii. These trees can similarly be called interior subalpine fir, i.e., A. bifolia × lasiocarpa. Both A. lasiocarpa and A. bifolia need comparative morphologic studies.</p><!--
+
|discussion=<p><i>Abies bifolia</i> has been—and by many workers still is—included in synonymy under <i>A. lasiocarpa</i> or A. subalpina since about 1890, and A. subalpina under <i>A. lasiocarpa</i> since about the 1920s. <i>Abies bifolia</i> is distinct from <i>A. lasiocarpa</i>, however, in chemical tests on wood (H.S. Fraser and E.P. Swan 1972), lack of crystals in the ray parenchyma (R.W. Kennedy et al. 1968), lack of lasiocarpenonol (J.F. Manville and A.S. Tracey 1989), and distinct terpene patterns (R.S. Hunt and E.von Rudloff 1979). <i>Abies bifolia</i> also tends to have slightly shorter and fewer prominently notched leaves than <i>A. lasiocarpa</i>. The two are clearly separated by the color of their periderm and by the shape of their basal bud scales. These firs may be more distinct than the pairs <i>A. balsamea</i> -- <i>A. fraseri</i> and <i>A. procera</i> -- <i>A. magnifica</i>. A north-south transect, however, from south central Yukon to northern Washington yielded introgressed trees possessing characteristics of both <i>A. lasiocarpa</i> and <i>A. bifolia</i>, recalling the interior spruce (Canadian Forestry Service 1983), which has characteristics of both <i>Picea glauca</i> and <i>P. engelmannii</i>. These trees can similarly be called interior subalpine fir, i.e., <i>A. bifolia</i> × lasiocarpa. Both <i>A. lasiocarpa</i> and <i>A. bifolia</i> need comparative morphologic studies.</p><!--
--><p>Isolated southern populations of Abies bifolia may also have unique characteristics. The taxonomy of corkbark fir, treated by some as A. lasiocarpa var. arizonica (Merriam) Lemmon, is uncertain. This taxon should probably be a segregate of A. bifolia, not A. lasiocarpa, a disposition that requires a thorough morphologic and chemical reappraisal, especially since the work of E.Zavarin et al. (1970) suggested that populations south of Wyoming may have unique terpene patterns. In north central Alberta, A. bifolia introgresses with A. balsamea (R.S. Hunt and E.von Rudloff 1974; E.H. Moss 1953).</p>
+
--><p>Isolated southern populations of <i>Abies bifolia</i> may also have unique characteristics. The taxonomy of corkbark fir, treated by some as <i>A. lasiocarpa</i> <i></i>var.<i> arizonica</i> (Merriam) Lemmon, is uncertain. This taxon should probably be a segregate of <i>A. bifolia</i>, not <i>A. lasiocarpa</i>, a disposition that requires a thorough morphologic and chemical reappraisal, especially since the work of E.Zavarin et al. (1970) suggested that populations south of Wyoming may have unique terpene patterns. In north central Alberta, <i>A. bifolia</i> introgresses with <i>A. balsamea</i> (R.S. Hunt and E.von Rudloff 1974; E.H. Moss 1953).</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Abies bifolia
 
name=Abies bifolia
|author=
 
 
|authority=A. Murray bis
 
|authority=A. Murray bis
 
|rank=species
 
|rank=species
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|publication title=Proc. Roy. Hort. Soc. London
 
|publication title=Proc. Roy. Hort. Soc. London
 
|publication year=1863
 
|publication year=1863
|special status=
+
|special status=Endemic
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_576.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_576.xml
 
|genus=Abies
 
|genus=Abies
 
|species=Abies bifolia
 
|species=Abies bifolia
|adaxial surface coloration=light green;bluish green
 
|adaxial surface pubescence=glaucous
 
|age architecture=furrowed
 
|apex shape=rounded;slightly notched;rounded
 
|bark architecture or pubescence=scaly
 
|bark architecture or pubescence or relief=smooth
 
|bark coloration=gray
 
|bark width=thin
 
|basal scale coating=not resinous;resinous
 
|basal scale length or size=long
 
|basal scale pubescence=glabrous
 
|basal scale shape=triangular;spatulate
 
|basal scale size or width=narrow
 
|body coloration=brown
 
|bract position=included
 
|branch fragility=stiff
 
|branch fragility or size=stout
 
|branch orientation=diverging
 
|bud coating=resinous
 
|bud coloration=brown
 
|bud prominence=exposed
 
|bud shape=globose
 
|bud size=small
 
|cotyledon count=3;6
 
|cross-section architecture=grooved
 
|cross-section odor=camphorlike
 
|cross-section prominence or shape=flat
 
|crown shape=spirelike
 
|leaf arrangement=spiraled
 
|leaf fragility=flexible
 
|leaf length=1.1cm;2.5cm
 
|leaf orientation=turned
 
|leaf width=1.25mm;1.5mm
 
|leaf-scar condition=fresh
 
|margin position=midway
 
|margin shape=entire;rarely crenate
 
|periderm coloration=light-brown
 
|pollination coloration=purplish
 
|resin canal position=median
 
|resin canal size=large
 
|row architecture=continuous
 
|row count=numerous;3;6
 
|scale length=1.5
 
|scale pubescence=pubescent
 
|scale width=2.5
 
|seed length=5mm;7mm
 
|seed width=2mm;3mm
 
|seed-cone architecture=sessile
 
|seed-cone coloration=dark purple-blue;grayish purple
 
|seed-cone length=5cm;10cm
 
|seed-cone shape=cylindric
 
|seed-cone width=3cm;3.5cm
 
|stomatal count=3;6
 
|tree some measurement=0m;30m
 
|trunk diameter=0m;0.45m
 
|twig arrangement=opposite;whorled
 
|twig coloration=light-brown;grayish
 
|twig pubescence=sparse
 
|wing coloration=grayish brown
 
|wing length=1.5 times as long as body
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Abies]]
 
-->[[Category:Treatment]][[Category:Abies]]

Latest revision as of 20:24, 5 November 2020

Trees to 30m; trunk to 0.45m diam.; crown spirelike. Bark gray, thin, smooth, with age somewhat furrowed and scaly (toward southern end of range bark is corky [corkbark fir]). Branches diverging from trunk at right angles, stout, stiff; twigs opposite to whorled, grayish, pubescence sparse, light brown; fresh leaf scars with light brown periderm. Buds exposed, brown, globose, small, resinous, apex rounded; basal scales long, narrow, isosceles triangular to spatulate, glabrous, resinous or not resinous, margins entire to rarely crenate, apex sharp-pointed or rounded. Leaves 1.1–2.5cm × 1.25–1.5mm, spiraled and turned upward, flexible; cross section flat, grooved adaxially, sometimes only slightly so; odor camphorlike; abaxial surface with 3–5 stomatal rows on each side of midrib; adaxial surface light green to bluish green, usually glaucous, with 3–6 stomatal rows at midleaf, rows usually continuous to leaf base, usually more numerous toward leaf apex; apex slightly notched to rounded; resin canals large, ± median, away from margins and midway between abaxial and adaxial epidermal layers. Pollen cones at pollination purplish. Seed cones cylindric, 5–10 × 3–3.5cm, dark purple-blue to grayish purple, sessile, apex rounded; scales ca. 1.5 × 2.5cm, densely pubescent; bracts included. Seeds 5–7 × 2–3mm, body brown; wing about 1.5 times as long as body, grayish brown; cotyledons 3–6.


Habitat: Continental, subalpine coniferous forests
Elevation: 600–3600m

Distribution

V2 576-distribution-map.gif

Alta., B.C., N.W.T., Yukon, Ariz., Colo., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo.

Discussion

Abies bifolia has been—and by many workers still is—included in synonymy under A. lasiocarpa or A. subalpina since about 1890, and A. subalpina under A. lasiocarpa since about the 1920s. Abies bifolia is distinct from A. lasiocarpa, however, in chemical tests on wood (H.S. Fraser and E.P. Swan 1972), lack of crystals in the ray parenchyma (R.W. Kennedy et al. 1968), lack of lasiocarpenonol (J.F. Manville and A.S. Tracey 1989), and distinct terpene patterns (R.S. Hunt and E.von Rudloff 1979). Abies bifolia also tends to have slightly shorter and fewer prominently notched leaves than A. lasiocarpa. The two are clearly separated by the color of their periderm and by the shape of their basal bud scales. These firs may be more distinct than the pairs A. balsamea -- A. fraseri and A. procera -- A. magnifica. A north-south transect, however, from south central Yukon to northern Washington yielded introgressed trees possessing characteristics of both A. lasiocarpa and A. bifolia, recalling the interior spruce (Canadian Forestry Service 1983), which has characteristics of both Picea glauca and P. engelmannii. These trees can similarly be called interior subalpine fir, i.e., A. bifolia × lasiocarpa. Both A. lasiocarpa and A. bifolia need comparative morphologic studies.

Isolated southern populations of Abies bifolia may also have unique characteristics. The taxonomy of corkbark fir, treated by some as A. lasiocarpa var. arizonica (Merriam) Lemmon, is uncertain. This taxon should probably be a segregate of A. bifolia, not A. lasiocarpa, a disposition that requires a thorough morphologic and chemical reappraisal, especially since the work of E.Zavarin et al. (1970) suggested that populations south of Wyoming may have unique terpene patterns. In north central Alberta, A. bifolia introgresses with A. balsamea (R.S. Hunt and E.von Rudloff 1974; E.H. Moss 1953).

Selected References

None.

Lower Taxa

None.
... more about "Abies bifolia"
Richard S. Hunt +
A. Murray bis +
Rocky Mountain alpine fir +, Rocky Mountain subalpine fir +  and corkbark fir +
Alta. +, B.C. +, N.W.T. +, Yukon +, Ariz. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +  and Wyo. +
600–3600m +
Continental, subalpine coniferous forests +
Proc. Roy. Hort. Soc. London +
Abies subalpina +
Abies bifolia +
species +